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1 Leaves strongly 5-ranked; leaf cells mammillose or more or less smooth; rhizoids smooth; operculum rostrate; peristome teeth apically fused.   Conostomum
+ Leaves typically not 5-ranked (seriately spiralled inPhilonotis seriata); leaf cells prorulose, mammillose or smooth; rhizoids papillose; operculum convex, conic or umbonate; peristome teeth, when present, apically free.   (2)
2 (1) Leaves 3-ranked; leaf cells covered by a striated cuticle (leaves appearing papillose in section).   Plagiopus
+ Leaves not in distinct ranks or rarely 5-ranked; leaf cells prorulose, mammillose or rarely smooth.   (3)
3 (2) Stems lacking a hyalodermis; epidermis prorulose; capsules irregularly wrinkled; western North America.   Anacolia
+ Stem hyalodermis present (sometimes weakly developed); epidermis not prorulose; capsules usually furrowed (irregularly wrinkled in Philonotis cernua);wide ranging species or eastern North America or Greenland.   (4)
4 (3) Leaves 2-stratose (at least along distal margins); fertile plants lacking a subfloral whorl of branches.   Bartramia
+ Leaves 1-stratose; fertile plants often with a subfloral whorl of branches.   Philonotis

Lower Taxa


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