| 1 |
Petaline limb obscure, with a hammer-shaped spur and minute lip; carpels 7--9 |
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210 A. polycarpum |
| + |
Petaline limb distinct, with a short or long spur and distinct lip; carpels usually 3--5. |
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(2) |
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| 2 (1) |
Stem erect, ascending, reclining, or trailing, not twining. |
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(3) |
| + |
Stem twining. |
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(24) |
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| 3 (2) |
Stem leafy, not scapose. |
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(4) |
| + |
Stem scapose to subscapose. |
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(5) |
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| 4 (4) |
Leaf blade palmately divided |
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Key 4 |
| + |
Leaf blade undivided |
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105 A. rhombifolium |
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| 5 (3) |
Leaf blade larger, 7--20(--30) cm wide. |
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(6) |
| + |
Leaf blade smaller, less than 7 cm wide. |
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(8) |
| |
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| 6 (5) |
Leaf lobes subdissected, ultimate lobules narrowly lanceolate to linear |
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39 A. duclouxii |
| + |
Leaf lobes less divided, ultimate lobules triangular to lanceolate or broadly lanceolate. |
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(7) |
| |
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| 7 (6) |
Pedicels spreading pubescent; leaf lobes contiguous or slightly overlapping |
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37 A. coriophyllum |
| + |
Pedicels appressed pubescent; leaf lobes separate |
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38 A. nagarum |
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| 8 (5) |
Leaf blade deeply 3-lobed near to base. |
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(9) |
| + |
Leaf blade medially 3-lobed. |
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(13) |
| |
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| 9 (8) |
Stem, pedicels, and sepals abaxially glabrous; upper sepal galeate, with a long beak |
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149 A. monanthum |
| + |
Stem, pedicels, and sepals abaxially pubescent; upper sepal navicular or falcate-navicular. |
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(10) |
| |
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| 10 (9) |
Cauline leaves 6--9; upper sepal falcate-navicular. |
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(11) |
| + |
Cauline leaves 1 or 2; upper sepal navicular. |
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(12) |
| |
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| 11 (10) |
Inflorescence 3--10-flowered; pedicels to 5 cm; carpels glabrous |
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36 A. sinchiangense |
| + |
Inflorescence ca. 20-flowered; pedicels to 3 cm; carpels pubescent |
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178 A. handelianum |
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| 12 (10) |
Petaline limb with both lip and spur |
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34 A. pulchellum |
| + |
Petaline limb strongly reduced to a small hooklike structure, without lip and spur |
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35 A. hamatipetalum |
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| 13 (8) |
Petaline lip divided into 2 linear lobes |
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28 A. rotundifolium |
| + |
Petaline lip divided into 2 ± orbicular lobes. |
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(14) |
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| 14 (13) |
Petaline spur erect. |
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(15) |
| + |
Petaline spur slightly incurved. |
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(16) |
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| 15 (14) |
Bracteoles linear, undivided |
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26 A. naviculare |
| + |
Bracteoles 3-fid |
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27 A. creagromorphum |
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| 16 (14) |
Inflorescence retrorse pubescent. |
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(17) |
| + |
Inflorescence spreading pubescent. |
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(20) |
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| 17 (16) |
Bracts leaflike; bracteoles 3-fid; petals pubescent, distinctly spurred, lip tonguelike, distinct |
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30 A. longipetiolatum |
| + |
Bracts linear; bracteoles linear, entire; petals glabrous, with slightly incurved or cup-shaped spur, lip distinct or not. |
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(18) |
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| 18 (17) |
Petaline spur slightly incurved, lip indistinct, claw straight at spur apex |
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23 A. tanguticum |
| + |
Petaline spur cup-shaped, lip short but distinct, claw straight or coiled at spur apex. |
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(19) |
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| 19 (18) |
Petaline claw strongly coiled at apex, lip entire, projecting forward |
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25 A. spiripetalum |
| + |
Petaline claw slightly incurved at apex, lip 2-lobed, reflexed |
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24 A. qinghaiense |
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| 20 (16) |
Inflorescence racemose, 2--5-flowered; leaf blade to 5 cm wide; upper sepal navicular-conical to conical |
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29 A. changianum |
| + |
Inflorescence corymbose, 1--3-flowered; leaf blade usually to 3 cm wide (A. ichangense to 5.4 cm); upper sepal navicular to navicular-conical. |
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(21) |
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| 21 (20) |
Bracteoles narrowly elliptic; carpels 3 |
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31 A. ichangense |
| + |
Bracteoles subulate; carpels 5. |
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(22) |
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| 22 (21) |
Sepals yellow to greenish white; leaf lobes shallowly lobed |
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32 A. iochanicum |
| + |
Sepals blue-purple; leaf lobes deeply lobed. |
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(23) |
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| 23 (22) |
Flowers larger, upper sepal 2--2.5 cm from base to beak |
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33 A. alpinonepalense |
| + |
Flowers smaller, upper sepal to 1.5 cm from base to beak |
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34 A. pulchellum |
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| 24 (2) |
Pedicels spreading pubescent. |
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(25) |
| + |
Pedicels retrorse and appressed pubescent or glabrous. |
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(30) |
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| 25 (24) |
Leaf blade deeply 3-lobed to 2--3.3 mm from base, lobes ± dissected, lobules narrowly lanceolate to linear-lanceolate. |
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(26) |
| + |
Leaf blade medially to shallowly 3-lobed; segment undivided or lobed, rarely dissected (A. volubile). |
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(27) |
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| 26 (25) |
Sepals white, upper sepal galeate, ca. 1.5 cm high; bracteoles villous; carpels 3 |
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108 A. weixiense |
| + |
Sepals blue-purple, upper sepal high galeate, 1.7--1.8 cm high; bracteoles glabrous; carpels 5 |
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121 A. episcopale |
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| 27 (25) |
Leaf central segment undivided or indistinctly 3-lobed, margin coarsely dentate |
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126 A. cannabifolium |
| + |
Leaf central segment 3-lobed or pinnatisect. |
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(28) |
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| 28 (27) |
Stem spreading pilose; petals pubescent; carpels glabrous |
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125 A. shensiense |
| + |
Stem glabrous or sparsely retrorse pubescent; petals glabrous; ovary pubescent. |
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(29) |
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| 29 (28) |
Rhizome connecting caudices shorter than 5 mm; central leaf segment 3-lobed, ultimate lobules triangular |
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128 A. sczukinii |
| + |
Rhizome connecting caudices up to 6 cm; central leaf segment subpinnately parted, ultimate lobules lanceolate to linear |
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129 A. volubile |
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| 30 (24) |
Bracteoles elliptic, obovate, spatulate, to leaflike and palmately lobed. |
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(31) |
| + |
Bracteoles linear to subulate, entire. |
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(34) |
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| 31 (30) |
Stem bulbiferous; bracteoles palmately lobed; petaline spur ca. 10 mm |
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111 A. bulbilliferum |
| + |
Stem not bulbiferous; bracteoles entire; petaline spur 1.5--3 mm. |
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(32) |
| |
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| 32 (31) |
Leaf blade deeply 3-lobed less than 5 mm from base |
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90 A. contortum |
| + |
Leaf blade medially 3-lobed more than 5 mm from base. |
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(33) |
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| 33 (32) |
Leaf blade 3-parted to more than 1 cm from base, central lobe 3-lobed |
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107 A. refracticarpum |
| + |
Leaf blade 3-parted to 5--7 mm from base, central lobe pinnatisect |
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110 A. tsaii |
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| 34 (30) |
Leaf blade deeply 3-lobed less than 5 mm from base. |
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(35) |
| + |
Leaf blade medially 3-lobed more than 5 mm from base. |
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(39) |
| |
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| 35 (34) |
Leaf central segment undivided, bluntly dentate at margin, lanceolate or rhombic-lanceolate |
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127 A. henryi |
| + |
Leaf central segment lobed or parted, rhombic or broadly rhombic. |
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(36) |
| |
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| 36 (35) |
Leaf central segment broadly rhombic, acute at apex |
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117 A. vilmorinianum |
| + |
Leaf central segment rhombic, acuminate or long acuminate at apex. |
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(37) |
| |
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| 37 (36) |
Leaf segments dissected; ultimate lobules lanceolate to linear |
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129 A. volubile |
| + |
Leaf segments lobed; ultimate lobules ovate or triangular. |
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(38) |
| |
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| 38 (37) |
Upper sepal galeate, with a distinct beak; petals and carpels pubescent |
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123 A. lioui |
| + |
Upper sepal high galeate, with an indistinct beak; petals and carpels glabrous |
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124 A. sungpanense |
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| 39 (34) |
Leaf blade parted to more than 6 mm above base. |
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(40) |
| + |
Leaf blade parted to 1.5--5 mm from base. |
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(45) |
| |
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| 40 (39) |
Upper sepal high galeate. |
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(41) |
| + |
Upper sepal galeate. |
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(42) |
| |
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| 41 (40) |
Leaf blade 3-parted slightly beyond middle, central lobe ovate-rhombic or trapezoid-rhombic, indistinctly 3-lobed; pedicels appressed pubescent or glabrous |
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106 A. hemsleyanum |
| + |
Leaf blade 3-parted nearly to base, central lobe rhombic, 3-parted or subpinnately divided; pedicels densely retrorse pubescent |
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116 A. austroyunnanense |
| |
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| 42 (40) |
Leaf central lobe ovate-rhombic or trapezoid, nearly undivided, several dentate above middle |
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109 A. crassicaule |
| + |
Leaf central lobe broadly ovate-rhombic, pinnately divided. |
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(43) |
| |
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| 43 (42) |
Inflorescence rachis sparsely retrorse pubescent; petals sparsely pubescent; leaf blade leathery |
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119 A. validinerve |
| + |
Inflorescence rachis glabrous; petals glabrous; leaf blade membranous. |
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(44) |
| |
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| 44 (43) |
Leaf central lobe cuneate-rhombic, narrowly cuneate at base; petaline claw geniculate at apex |
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115 A. longilobum |
| + |
Leaf central lobe rhombic, broadly cuneate at base; petaline claw not geniculate |
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118 A. pseudostapfianum |
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| 45 (39) |
Upper sepal high galeate. |
|
(46) |
| + |
Upper sepal galeate to conical or hemispheric. |
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(48) |
| |
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|
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| 46 (45) |
Leaf lobes lobed, ultimate lobules triangular or narrowly triangular |
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116 A. austroyunnanense |
| + |
Leaf lobes parted, ultimate lobules sublinear. |
|
(47) |
| |
|
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| 47 (46) |
Carpels glabrous; inflorescence 2--4-flowered |
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114 A. elwesii |
| + |
Carpels sparsely appressed pubescent; inflorescence 10--16-flowered |
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122 A. campylorrhynchum |
| |
|
|
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| 48 (45) |
Pedicels and sepals abaxially glabrous |
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120 A. stapfianum |
| + |
Pedicels and sepals abaxially pubescent. |
|
(49) |
| |
|
|
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| 49 (48) |
Upper sepal with a beak ca. 5 mm, usually curved upward |
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122 A. campylorrhynchum |
| + |
Upper sepal with a beak ca. 0.5 mm, straight or curved downward. |
|
(50) |
| |
|
|
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| 50 (49) |
Petaline claw not geniculate at apex; pedicels 7--26 mm; leaf central lobe 3-lobed below middle |
|
113 A. chuanum |
| + |
Petaline claw geniculate at apex; pedicels 22--70 mm; leaf central lobe 3-parted beyond middle |
|
112 A. habaense |
