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FOC | Family List | FOC Vol. 6 | Ranunculaceae

10. Aconitum Linnaeus, Sp. Pl. 1: 532. 1753; Gen. Pl. ed. 5, 236; 1754.

乌头属 wu tou shu

Li Liangqian; Yuichi Kadota

Herbs perennial or pseudoannual, rarely annual, with taproots or 2 to several caudices. Stem erect or twining. Leaves simple or compound, cauline ones alternate, sometimes all basal, palmately divided, rarely undivided. Inflorescence usually racemose. Pedicel with 2 bracteoles. Flowers bisexual, zygomorphic. Sepals 5, petaloid, purple, blue, or yellow; lower sepals 2, narrowly lanceolate or oblong, small; lateral sepals 2, suborbicular; upper sepal falcate, navicular, galeate to cylindric. Petals 2, clawed; limb usually with lip and spur, secretory tissue usually at limb apex, rarely abaxial. Staminodes usually absent. Stamens numerous; anthers ellipsoid-globose. Carpels 3--5(--13); style short, persistent.

About 400 species: temperate regions of the N hemisphere; 211 species (166 endemic) in China.


Key 4

1 Leaf blade medially to shallowly 3-lobed, lobes usually dissected.   (2)
+ Leaf blade ternate to deeply 3-lobed.   (59)
       
2 (1) Central lobe of middle cauline leaves undivided or indistinctly 3-lobed, lateral lobes undivided or unequally 2-lobed, distal leaves undivided, rhombic-ovate; upper sepal subcylindric   104 A. racemulosum
+ Central and lateral lobes of cauline leaves distinctly divided; upper sepal high galeate or galeate to falcate.   (3)
       
3 (2) Lateral sepals 6.5--8.5 mm in diam.   (4)
+ Lateral sepals more than 10 cm.   (7)
       
4 (3) Flower solitary, axillary; bracteoles leaflike, 1.5--1.8 cm; upper sepal navicular-conical   46 A. sinoaxillare
+ Flowers in an inflorescence; bracteoles linear to lanceolate-linear, or subulate, 1.5--4 mm; upper sepal falcate-navicular or conical.   (5)
       
5 (4) Inflorescence ca. 5 cm, 4-flowered; petals shortly spurred, claw geniculate at apex; leaf central lobe rhombic   48 A. milinense
+ Inflorescence 14--50 cm, 15--30-flowered; petals not spurred, claw not geniculate at apex; leaf central lobe rhombic-obovate to truncate-rhombic.   (6)
       
6 (5) Stem ca. 50 cm tall; leaf blade 3.2--3.5 × 5--7 cm; upper sepal navicular, not contracted   49 A. brunneum
+ Stem 85--100 cm tall; leaf blade 3.8--6 × 6.5--11 cm; upper sepal galeate, contracted   50 A. pseudobrunneum
       
7 (3) Inflorescence rachis, and pedicels glabrous or only rachis retrorse pubescent.   (8)
+ Inflorescence rachis and pedicels pubescent, in A. shimianense pedicels glabrous and in A. lihsienense rachis glabrous.   (20)
       
8 (7) Inflorescence rachis retrorse pubescent; pedicels and carpels glabrous   59 A. shimianense
+ Inflorescence rachis, pedicels, and carpels glabrous or pedicels nearly glabrous in A. fanjingshanicum, slightly spreading pubescent at apex.   (9)
       
9 (8) Bracteoles leaflike, 3-lobed   58 A. pukeense
+ Bracteoles entire.   (10)
       
10 (9) Bracteoles narrowly obovate or spatulate   52 A. spathulatum
+ Bracteoles linear, elliptic, or subulate.   (11)
       
11 (10) Stem trailing distally, distally S-curved; inflorescence unequally bifurcate-branched   130 A. fischeri
+ Stem erect, distally not S-curved; inflorescence not bifurcate-branched.   (12)
       
12 (11) Bracteoles adaxially pubescent and ciliate; pedicels nearly glabrous   92 A. fanjingshanicum
+ Bracteoles and pedicels glabrous.   (13)
       
13 (12) Petaline claw geniculate at apex.   (14)
+ Petaline claw not geniculate at apex.   (15)
       
14 (13) Bracteoles at middle of pedicels, linear, 3.5--4.5 mm   53 A. geniculatum
+ Bracteoles contiguous with flower, leaflike, 1--3.5 cm   55 A. magnibracteolatum
       
15 (13) Upper sepal navicular, 2.5--3 cm from base to beak   70 A. yunlingense
+ Upper sepal galeate to high galeate, less than 2.1 cm from base to beak.   (16)
       
16 (15) Leaf blade adaxially appressed pubescent, abaxially sparsely pubescent at veins, petioles sparsely spreading pubescent   54 A. luningense
+ Leaf blade adaxially sparsely pubescent, glabrous, or sparsely retrorse pubescent, abaxially and petioles glabrous.   (17)
       
17 (16) Upper sepal high galeate, lower margin straight   91 A. lonchodontum
+ Upper sepal galeate to conical, lower margin oblique.   (18)
       
18 (17) Leaf blade leathery; lobes usually slightly overlapping; carpels 3   68 A. coriaceifolium
+ Leaf blade membranous; lobes separate; carpels 5.   (19)
       
19 (18) Leaf blade 9--11 × 13--18 cm; petaline spur incurved, ca. 1.8 mm; carpels 5   51 A. bulleyanum
+ Leaf blade ca. 2.7 × 4.5 cm; petaline spur circinate, ca. 7 mm; carpels 4   79 A. namlaense
       
20 (7) Inflorescence rachis and pedicels retrorse pubescent, in A. shimianense pedicels glabrous.   (21)
+ Pedicels spreading pubescent.   (35)
       
21 (20) Pedicels glabrous   59 A. shimianense
+ Pedicels retrorse pubescent.   (22)
       
22 (21) Upper sepal high galeate or subcylindric.   (23)
+ Upper sepal navicular to galeate.   (27)
       
23 (22) Bracteoles of lower pedicels leaflike; petaline spur sparsely pubescent   152 A. jiulongense
+ Bracteoles not leaflike; petaline spur usually glabrous.   (24)
       
24 (23) Leaf central lobe narrowly rhombic, narrowly cuneate at base; upper sepal subcylindric; filaments puberulous   130 A. fischeri
+ Leaf central lobe rhombic, broadly cuneate at base; upper sepal high galeate; filaments glabrous.   (25)
       
25 (24) Bracteoles sparsely pubescent; carpels 3, densely appressed pubescent   56 A. yangii
+ Bracteoles glabrous; carpels 3(--5), pubescent.   (26)
       
26 (25) Leaf lobules slightly obtuse at apex; inflorescence loosely ca. 10-flowered; upper sepal with beak to 7 mm   57 A. georgei
+ Leaf lobules acute at apex; inflorescence densely many flowered; upper sepal with indistinct beak 1--2 mm   135 A. carmichaelii
       
27 (22) Upper sepal navicular.   (28)
+ Upper sepal galeate.   (29)
       
28 (27) Pedicels ascending at an obtuse angle; upper sepal broadest above middle, ca. 2.5 cm from base to beak, lower margin straight   77 A. brevilimbum
+ Pedicels erect or ascending at an acute angle; upper sepal broadest at middle, ca. 1.9 cm from base to beak, lower margin concave   96 A. longipedicellatum
       
29 (27) Inflorescence corymbose, 2--4-flowered   93 A. taipeicum
+ Inflorescence racemose or paniculate, usually many flowered.   (30)
       
30 (29) Pedicels incurved above   87 A. pendulicarpum
+ Pedicels ascending at an obtuse angle.   (31)
       
31 (30) Sepals yellowish; bracteoles pubescent; petaline spur circinate   85 A. alboflavidum
+ Sepals blue; bracteoles glabrous; petaline spur not circinate.   (32)
       
32 (31) Stem 40--50 cm tall; central leaf lobe slightly obtuse at apex; inflorescence 4- or 5-flowered; upper sepal ca. 1.4 cm from base to beak   78 A. chiachaense
+ Stem 100--120 cm tall; central leaf lobe acuminate or long acuminate at apex; inflorescence 2--10-flowered; upper sepal 1.9--2.3 cm from base to beak.   (33)
       
33 (32) Upper sepal abaxially and carpels pubescent   94 A. transsectum
+ Upper sepal abaxially and carpels glabrous.   (34)
       
34 (33) Leaf blade nearly glabrous or sparsely pubescent on both surfaces; bracteoles of lower pedicels 3-lobed; petals glabrous   64 A. franchetii
+ Leaf blade adaxially sparsely appressed puberulous, abaxially retrorse puberulous at veins; bracteoles linear; petals sparsely pubescent   83 A. lobulatum
       
35 (20) Upper sepal yellow or green-tinged.   (36)
+ Upper sepal blue-purple, rarely white.   (38)
       
36 (35) Inflorescence rachis glabrous; pedicels spreading white pubescent   84 A. stramineiflorum
+ Inflorescence rachis densely spreading yellow pubescent.   (37)
       
37 (36) Bracteoles proximally on pedicels; upper sepal navicular-galeate or galeate; carpels densely pubescent; stem 25--70 cm tall   80 A. souliei
+ Bracteoles distally on pedicels; upper sepal navicular; carpels glabrous; stem ca. 30 cm tall   82 A. phyllostegium
       
38 (35) Upper sepal navicular or navicular-galeate.   (39)
+ Upper sepal galeate or high galeate.   (48)
       
39 (38) Bracteoles leaflike, 3-parted.   (40)
+ Bracteoles undivided, oblong, oblanceolate, linear, or subulate.   (42)
       
40 (39) Petals and carpels glabrous; petaline claw geniculate at apex   60 A. pomeense
+ Petals and carpels pubescent; petaline claw not geniculate at apex.   (41)
       
41 (40) Lower pedicels 6--12 cm; upper sepal abaxially nearly glabrous, with a beak, lateral sepals 1.7--2.3 cm   69 A. stylosum
+ Lower pedicels 2.5--6.5 cm; upper sepal abaxially densely pubescent; lateral sepals 1.5--1.7 cm with blunt beak   63 A. nielamuense
       
42 (39) Pedicels incurved; sepals abaxially and carpels both glabrous.   (43)
+ Pedicels straight; sepals abaxially or at margin and carpels all pubescent.   (44)
       
43 (42) Stem erect, long branched; branches to 18 cm; central leaf lobe acuminate; petals glabrous, claw geniculate at apex   86 A. nutantiflorum
+ Stem trailing, branched; branches to 35 cm; central leaf lobe caudate-acuminate; petals pubescent, claw not geniculate at apex   103 A. longiramosum
       
44 (42) Stem 30--35 cm tall; leaf blade adaxially glabrous, abaxially pubescent at veins   81 A. parcifolium
+ Stem taller than 70 cm ; leaf blade pubescent on both surfaces or abaxially subglabrous.   (45)
       
45 (44) Stem and inflorescence rachis glabrous   95 A. lihsienense
+ Stem and inflorescence rachis pubescent.   (46)
       
46 (45) Inflorescence ca. 7-flowered, rachis white pubescent; upper sepal ca. 2.5 cm from base to beak   71 A. stylosoides
+ Inflorescence densely many flowered, rachis yellow pubescent; upper sepal 1.2--2.1 cm.   (47)
       
47 (46) Petals glabrous; carpels 5; petioles 3--7 cm; upper sepal ca. 1.2 cm from base to beak   73 A. kungshanense
+ Petals sparsely pubescent; carpels 3; petioles 7--15 cm; upper sepal ca. 2.1 cm from base to beak   66 A. lycoctonifolium
       
48 (38) Upper sepal high galeate   151 A. piepunense
+ Upper sepal galeate.   (49)
       
49 (48) Middle cauline leaf shortly petiolate, upper ones subsessile   67 A. forrestii
+ Middle cauline leaf with petioles nearly as long as leaf blade.   (50)
       
50 (49) Filaments pubescent.   (51)
+ Filaments glabrous.   (53)
       
51 (50) Leaf central lobe rhombic or broadly rhombic, lobed, acute or shortly acuminate at apex; petaline claw sparsely puberulous   76 A. spicatum
+ Leaf central lobe narrowly rhombic or rhombic, parted, acuminate or long acuminate at apex; petaline claw densely pubescent.   (52)
       
52 (51) Bracteoles proximally on pedicels, linear or linear-subulate; stem 60--170 cm tall, inflorescence 7--30-flowered   74 A. tatsienense
+ Bracteoles contiguous with flower, leaflike or narrowly elliptic; stem 32--42 cm tall; inflorescence 3--5-flowered   65 A. secundiflorum
       
53 (50) Carpels densely glabrous.   (54)
+ Carpels densely pubescent.   (56)
       
54 (53) Inflorescence rachis apically and pedicels densely spreading white pubescent   89 A. dolichorhynchum
+ Inflorescence rachis and pedicels spreading yellowish or spreading yellow pubescent (in A. rockii appressed pubescent).   (55)
       
55 (54) Inflorescence rachis and pedicels spreading pubescent; upper sepal 2.5--3 cm from base to beak   75 A. elliotii
+ Inflorescence rachis and pedicels both spreading and appressed pubescent; upper sepal to 2 cm from base to beak   88 A. rockii
       
56 (53) Bracteoles leaflike.   (57)
+ Bracteoles linear (in A. taronense bracteoles of lowermost pedicels 3-fid).   (58)
       
57 (56) Stem and inflorescence rachis and pedicels densely yellowish pubescent; leaf central lobe rhombic, acuminate at apex; leaf blade pubescent on both surfaces; sepals white   61 A. yachiangense
+ Stem apically and inflorescence rachis glabrous, pedicels whitish pubescent; leaf central lobe broadly rhombic, acute at apex; leaf blade adaxially glabrous; sepals brown-purple   62 A. chayuense
       
58 (56) Leaf central lobe rhombic-obovate, 3-lobed, obtuse at apex, lobules obtuse at apex; upper sepal abaxially pubescent   72 A. taronense
+ Leaf central lobe rhombic, subpinnately parted, acuminate at apex, lobules slightly acute at apex; upper sepal abaxially glabrous   88 A. rockii
       
59 (1) Leaf lobe serrate to shallowly divided, ultimate lobules ovate or narrowly ovate.   (60)
+ Leaf lobes deeply divided, ultimate lobules narrowly lanceolate to linear.   (86)
       
60 (59) Stem trailing, long branched, glabrous; inflorescence unequally dichotomous branching, glabrous; bracteoles leaflike, 3-lobed   131 A. raddeanum
+ Stem erect or sometimes ascending, usually shortly branched; inflorescence rachis straight (in A. dunhuaense slightly dichotomously branched); bracteoles narrowly oblong or linear, entire.   (61)
       
61 (61) Caudices several, in a chain   Key 6
+ Caudices 2 or more, not in a chain.   (62)
       
62 (61) Leaf segments acuminate at apex (Taiwan).   (63)
+ Leaf segments acute at apex (mainland China).   (65)
       
63 (62) Stem and pedicels retrorse pubescent   133 A. kojimae
+ Stem and pedicels spreading pubescent.   (64)
       
64 (63) Stem erect, less branched; upper sepal navicular; follicles erect   132 A. fukutomei
+ Stem twining, much branched; upper sepal hemispheric; follicles divergent   134 A. formosanum
       
65 (62) Sepals yellowish or white.   (66)
+ Sepals purple-blue.   (67)
       
66 (65) Sepals yellowish; petals, stamens, and 5 carpels glabrous   137 A. wuchagouense
+ Sepals white; petals, stamens, and 3 carpels pubescent   174 A. xiangchengense
       
67 (65) Pedicels glabrous.   (68)
+ Pedicels pubescent.   (70)
       
68 (67) Stem twining; upper sepal pyramidal; bracteoles broadly oblong to 3-lobed   90 A. contortum
+ Stem erect; upper sepal galeate; bracteoles linear or subulate.   (69)
       
69 (68) Pedicels to 5 cm, straight upward; flowers ± crowded   136 A. kusnezoffii
+ Pedicels to 8 cm, oblique upward; flowers sparsely arranged   138 A. birobidshanicum
       
70 (67) Pedicels appressed pubescent.   (71)
+ Pedicels spreading pubescent.   (76)
       
71 (70) Leaf central lobe acute at apex.   (72)
+ Leaf central lobe long acuminate at apex.   (74)
       
72 (71) Leaf blade deeply 3-lobed, not ternate; upper sepal high galeate, 2--2.6 cm high   135 A. carmichaelii
+ Leaf blade ternate; upper sepal hemispheric or galeate, 1--2.5 cm high.   (73)
       
73 (72) Upper sepal hemispheric, ca. 1 cm high; petaline claw geniculate at apex   101 A. yanyuanense
+ Upper sepal galeate, 1--2.5 cm high; petaline claw not geniculate at apex   102 A. japonicum
       
74 (71) Inflorescence rachis glabrous; pedicels appressed puberulous only at apex   136 A. kusnezoffii
+ Inflorescence rachis and pedicels densely appressed pubescent.   (75)
       
75 (74) Leaf blade sparsely appressed pubescent on both surfaces; bracteoles 4--5 mm; petals sparsely pubescent; carpels 4   98 A. chuosjiaense
+ Leaf blade abaxially glabrous; bracteoles 8--15 cm; petals glabrous; carpels 5   139 A. bailangense
       
76 (70) Inflorescence rachis glabrous; pedicels spreading pubescent only near apex.   (77)
+ Inflorescence rachis and pedicels spreading or retrorse pubescent.   (79)
       
77 (76) Inflorescence dichotomously branched; pedicels 5--8.5 cm   142 A. dunhuaense
+ Inflorescence not dichotomously branched; pedicels less than 5 cm.   (78)
       
78 (77) Leaf lobes coarsely dentate; petaline lip slightly emarginate   140 A. jaluense
+ Leaf lobes pinnatisect; petaline lip 2-lobed at apex   141 A. paniculigerum
       
79 (76) Bracteoles ovate or broadly ovate.   (80)
+ Bracteoles linear.   (81)
       
80 (79) Stem and leaf blade glabrous; petals glabrous; pedicels 2.5--3.5 cm   40 A. hicksii
+ Stem and leaf blade pubescent; petals sparsely villous; pedicels 1--2.3 cm   41 A. jilongense
       
81 (79) Upper sepal galeate; petals puberulous.   (82)
+ Upper sepal high galeate; petals glabrous.   (84)
       
82 (81) Sepals abaxially glabrous; carpels pubescent   100 A. pseudodivaricatum
+ Sepals abaxially pubescent; carpels glabrous.   (83)
       
83 (82) Inflorescence rachis, pedicels, and leaf blade yellowish pubescent   99 A. zhaojiueense
+ Inflorescence rachis, pedicels, and leaf blade whitish pubescent   154 A. legendrei
       
84 (81) Leaf central segment broadly rhombic, acute at apex   135 A. carmichaelii
+ Leaf central segment rhombic or narrowly rhombic, acuminate or long acuminate at apex.   (85)
       
85 (84) Leaf central lobes coarsely dentate   140 A. jaluense
+ Leaf central lobes pinnatisect   141 A. paniculigerum
       
86 (59) Middle and distal leaves aggregated, very shortly petiolate or sessile.   (87)
+ Middle and distal leaves sparsely arranged, middle cauline leaves petiolate, petioles subequaling leaf blades, upper petioles becoming short.   (97)
       
87 (87) Caudices several forming a chain   Key 6
+ Caudices 2, rarely more and then not forming a chain.   (88)
       
88 (87) Inflorescence and sepals abaxially glabrous.   (89)
+ Inflorescence and sepals abaxially pubescent.   (90)
       
89 (88) Sepals purple-blue   201 A. glabrisepalum
+ Sepals yellowish   206 A. chilienshanicum
       
90 (88) Inflorescence spreading pubescent; upper sepal falcate-navicular.   (91)
+ Inflorescence appressed pubescent; upper sepal navicular, low galeate, or hemispheric.   (92)
       
91 (90) Sepals yellowish or blue-purple   205 A. pendulum
+ Sepals blue-purple   207 A. ludlowii
       
92 (90) Bracteoles contiguous with flower, leaflike or narrowly oblong   200 A. sessiliflorum
+ Bracteoles remote from flower, linear or subulate, rarely leaflike or broadly linear.   (93)
       
93 (92) Carpels 3.   (94)
+ Carpels 5.   (95)
       
94 (93) Lower pedicels ascending, to 3 cm; upper sepal navicular or low galeate, 1.5--1.7 cm from base to beak   202 A. polyschistum
+ Lower pedicels erect, ca. 3 mm; upper sepal hemispheric, ca. 2 cm from base to beak   199 A. brachypodum
       
95 (93) Bracteoles usually 3-lobed, broadly linear, 5--9 × ca. 1.5 mm; carpels ± spreading densely yellowish villous   199 A. brachypodum
+ Bracteoles undivided, subulate, 3--6 × ca. 0.5 mm; carpels sparsely appressed puberulous.   (96)
       
96 (95) Leaf blade glabrous; sepals yellowish green or dark purple, abaxially pubescent; petals sparsely pubescent   203 A. flavum
+ Leaf blade adaxially retrorse pubescent; sepals blue-purple, abaxially retrorse pubescent; petals glabrous   204 A. yinschanicum
       
97 (86) Inflorescence rachis and pedicels pubescent (in A. fusungense and A. laevicaule rachis glabrous).   (98)
+ Inflorescence rachis and pedicels glabrous throughout.   (107)
       
98 (97) Lateral sepals broadly triangular apically, acute or slightly acute at apex.   (99)
+ Lateral sepals orbicular-obovate, rounded or truncate at apex.   (101)
       
99 (98) Sepals greenish white, upper sepal ca. 1.4 cm from base to beak; petals ca. 6 mm, not spurred; carpels 3, glabrous   177 A. brevipetalum
+ Sepals blue-purple, upper sepal 1.6--1.8 cm from base to beak; petals 15--19 cm, spur incurved; carpels 5, ovary pubescent.   (100)
       
100 (99) Stem 75--120 cm tall; leaf blade 8--12 cm wide; inflorescence many flowered; bracteoles linear   172 A. acutiusculum
+ Stem ca. 30 cm tall; leaf blade to 4.5 cm wide; inflorescence ca. 10-flowered; bracteoles narrowly triangular   188 A. falciforme
       
101 (98) Leaf lobules acute or acuminate at apex (in A. smithii slightly obtuse at apex); petaline spur incurved.   (102)
+ Leaf lobules rounded, obtuse, or slightly acute at apex; petaline spur not incurved.   (104)
       
102 (101) Bracteoles broadly ovate or orbicular   42 A. bracteolatum
+ Bracteoles linear or subulate (in A. incisofidum leaflike).   (103)
       
103 (103) Caudices 2 (if several, then not forming a chain)   Key 5
+ Caudices to 10 forming a chain   Key 6
       
104 (101) Ovary glabrous   47 A. jinyangense
+ Ovary densely appressed or spreading pubescent.   (105)
       
105 (104) Inflorescence 2- or 3-flowered; bracteoles lanceolate-linear or narrowly linear; petaline limb ca. 4 mm, lip ca. 1.8 mm   44 A. wolongense
+ Inflorescence 8--12-flowered; bracteoles obovate or broadly elliptic or narrowly elliptic; petaline limb 6--8 mm, lip 4--5 mm.   (106)
       
106 (105) Inflorescence retrorse pubescent; bracteoles obovate or broadly elliptic; filaments puberulous   45 A. pseudohuiliense
+ Inflorescence spreading pubescent; bracteoles narrowly elliptic; filaments glabrous   43 A. huiliense
       
107 (97) Upper sepal navicular or navicular-galeate.   (108)
+ Upper sepal galeate.   (112)
       
108 (107) Bracteoles lobed or at least those of lower pedicels leaflike.   (109)
+ Bracteoles undivided, linear or subulate; upper sepal navicular.   (110)
       
109 (108) Stem basally pubescent; upper sepal navicular-galeate; petaline limb pubescent, claw not geniculate at apex   167 A. gammiei
+ Stem glabrous; upper sepal falcate-navicular; petals glabrous, claw geniculate at apex   181 A. tangense
       
110 (108) Bracteoles near base of pedicels; filaments sparsely puberulous at apex   189 A. maowenense
+ Bracteoles at middle or distal part of pedicels; filaments glabrous.   (111)
       
111 (110) Stem ca. 70 cm tall; leaf lobules narrowly lanceolate or lanceolate-linear; inflorescence ca. 30-flowered; petaline claw geniculate at apex; carpels 3--5   182 A. fangianum
+ Stem 20--50 cm tall; leaf lobules linear or narrowly linear; inflorescence 2--7-flowered; petaline claw not geniculate; carpels 3   195 A. jeholense
       
112 (107) Petaline limb ca. 1 mm wide   159 A. ambiguum
+ Petaline limb more than 2 mm wide.   (113)
       
113 (112) Inflorescence 1- or 2- to several flowered, corymbose   149 A. monanthum
+ Inflorescence racemose, more than 7-flowered.   (114)
       
114 (113) Sepals abaxially sparsely pubescent   144 A. ningwuense
+ Sepal abaxially glabrous.   (115)
       
115 (114) Upper sepal 1.2--1.4 cm high; petaline claw geniculate at apex.   (116)
+ Upper sepal 1.8--2 cm high; petaline claw not geniculate at apex.   (117)
       
116 (115) Stem glabrous; terminal inflorescence ca. 35 cm, ca. 40-flowered; petals glabrous, spur ca. 1 mm, subglobose   164 A. leiostachyum
+ Stem pubescent; terminal inflorescence ca. 7 cm, 7-flowered; petaline lip puberulous, spur ca. 5 mm, subcylindric   165 A. pseudogeniculatum
       
117 (115) Leaf ultimate lobules triangular to lanceolate; carpels 5   136 A. kusnezoffii
+ Leaf ultimate lobules lanceolate to linear; carpels 3   194 A. soongoricum

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