1 |
Involucre 8-12(-15) mm, spherical or broadly campanulate; capitula usually few, arranged in sparse to compound corymbiform synflorescences, or sometimes solitary |
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(2) |
+ |
Involucre 4-8 mm, obovate, campanulate, or semispherical; phyllaries obtuse or rounded at apex, rarely acuminate, usually expanding after flowering; capitula usually numerous, rarely few, congested to compound corymbiform or corymbiform at apex of stems or branches |
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(11) |
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2 (1) |
Leaves not decurrent on stems; involucre spherical, capitula usually lax |
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(3) |
+ |
Leaves somewhat decurrent and winglike on stems; capitula densely congested forming globose corymbs, or solitary |
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(6) |
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3 (2) |
Rhizome thickish; cauline middle leaves linear-lanceolate, cinereous or yellowish lanate; capitula congested, densely corymbiform. |
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54 A. acutifolia |
+ |
Rhizome slender, sarmentose; cauline leaves oblong, oblanceolate, or spatulate; capitula usually lax corymbiform or solitary |
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(4) |
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4 (3) |
Phyllaries white or yellowish white, base reddish brown; both surfaces of leaves gray lanate; Taiwan. |
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53 A. nagasawae |
+ |
Phyllaries white, base dark brown; leaves adaxially arachnoid, abaxially densely gray lanate |
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(5) |
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5 (4) |
Stems slender, lower parts 1-2 mm in diam.; leaf base not obviously amplexicaul, 3-veined, occasionally 1-veined; capitula few (or many) or solitary. |
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51 A. nepalensis |
+ |
Stems thickish, lower parts up to 4 mm in diam.; leaves broad and large, base amplexicaul, 3- or 5-veined. |
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52 A. triplinervis |
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6 (2) |
Perennial herbs, rhizome slender; involucre broadly campanulate or semispherical, 8-11 mm |
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(7) |
+ |
Small or pulvinate subshrubs; involucre spherical |
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(8) |
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7 (6) |
Phyllaries yellowish white, yellow, or sulfureous, rarely reddish; leaves gray or yellowish white arachnoid tomentose on both surfaces. |
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49 A. flavescens |
+ |
Phyllaries white; leaves somewhat green, both surfaces with capitate glandular hairs and canescent arachnoid tomentum on margin. |
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50 A. hancockii |
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8 (6) |
Stems and both surfaces of leaves grayish, tangled-cottony, felty-papery tomentose. |
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45 A. chlamydophylla |
+ |
Stems and both surfaces of leaves usually without above indumentum |
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(9) |
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9 (8) |
Stems and both surfaces of leaves yellowish brown long lanate; phyllaries wax-yellow or dark yellow; stems 6-20 cm. |
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46 A. pannosa |
+ |
Stems and both surfaces of leaves yellowish or grayish white lanate; stems 3-8 cm or acaulescent |
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(10) |
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10 (9) |
Both surfaces of leaves yellowish green, grayish white, or yellowish white tomentose; phyllaries white or yellowish white, base brown. |
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48 A. viridis |
+ |
Both surfaces of leaves cinereous or grayish white lanate; phyllaries white, base dark brown. |
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47 A. cinerascens |
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11 (1) |
Leaves not or scarcely decurrent on stems |
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(12) |
+ |
Leaves decurrent to broadly or narrowly winged on stems |
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(21) |
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12 (11) |
Leaf base cordate or auriculate, semiamplexicaul, margin recurved, thin or submembranous |
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(13) |
+ |
Leaf base attenuate, not obviously amplexicaul, margin flat or recurved, thick |
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(14) |
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13 (12) |
Plants tall, stems thickish, lower parts 2-4 mm in diam.; leaves large, 1.5-6 cm, arachnoid or glabrous adaxially. |
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8 A. contorta |
+ |
Plants short, stems slender, lower parts ca. 1 mm in diam.; leaves 0.5-1.5 cm, lanate on both surfaces. |
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9 A. hondae |
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14 (12) |
Multibranched subshrubs, usually fastigiate; leaves narrowly linear, margin recurved, densely white tomentose on both surfaces. |
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10 A. virgata |
+ |
Perennial herbs, or stem base woody, or subshrubs, unbranched or branched but not fastigiate; leaves not linear, margin flat or recurved |
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(15) |
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15 (14) |
Leaves gradually larger upward from middle, densely arranged to radiate under synflorescences, usually longer than compound corymb or corymb |
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(16) |
+ |
Leaves gradually smaller upward from middle, not dense under synflorescences, upper ones shorter than compound corymb or corymb |
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(17) |
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16 (15) |
Upper leaves elliptic, apex acuminate; phyllaries white. |
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32 A. oxyphylla |
+ |
Upper leaves oblong-lanceolate, apex acute; phyllaries purple-red or margin white. |
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33 A. porphyrolepis |
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17 (15) |
Stems gray tangled papery tomentose; leaves white cottony tomentose. |
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7 A. contortiformis |
+ |
Stems gray tomentose; leaves arachnoid or later glabrous adaxially, densely gray to rufous tomentose abaxially, or both surfaces white tomentose |
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(18) |
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18 (17) |
Leaves densely white tomentose, 0.8-1 cm, margin recurved; middle phyllaries with longitudinal corrugation, apex rounded. |
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6 A. plicata |
+ |
Leaves arachnoid or later glabrous adaxially, densely gray to rufous tomentose abaxially, 1.5-10 cm; middle phyllaries without longitudinal corrugation |
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(19) |
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19 (18) |
Leaves linear or linear-lanceolate, 5-10 cm, margin flat, base not decurrent on stems; phyllaries milky white, apex obtuse or rounded, rarely acuminate. |
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3 A. margaritacea |
+ |
Leaves usually linear or linear-oblong, 1.5-3.5 cm, base slightly decurrent on stems; phyllaries snow white or yellowish white |
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(20) |
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20 (19) |
Involucre 5-7 mm, phyllaries snow white; Xizang. |
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4 A. royleana |
+ |
Involucre ca. 5 mm, phyllaries yellowish white; Taiwan. |
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5 A. morrisonicola |
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21 (11) |
Stems, leaves, and involucre base with rust-colored stalked adhesive hairs; involucre obovate, phyllaries beige, membranous, translucent, not expanding. |
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1 A. bulleyana |
+ |
Stems, leaves, and involucre base gray, white, or yellowish brown lanate, with squarrose or capitate-stalked glandular hairs, or arachnoid, or somewhat glabrous; involucre campanulate or semispherical, phyllaries white or reddish, usually scarious, opaque, rarely membranous, somewhat expanding |
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(22) |
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22 (21) |
Biennial herbs with thickish taproot. |
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2 A. busua |
+ |
Perennial herbs with rhizome, or multibranched subshrubs or small subshrubs |
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(23) |
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23 (22) |
Both surfaces of leaves arachnoid at first, later glabrous except midvein and margin, without glandular hairs, dark green when dry; perennial herbs. |
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11 A. delavayi |
+ |
Both surfaces of leaves or only abaxially tomentose, or both surfaces or adaxially with squarrose or capitate-stalked glandular hairs, or adaxially glabrous and green when dry; subshrubs or perennial herbs |
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(24) |
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24 (23) |
Subshrubs or small subshrubs, stems multibranched |
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(25) |
+ |
Perennial herbs with rhizome, sometimes with taproot, stems unbranched or lower part woody and with developing axillary buds or short branches |
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(29) |
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25 (24) |
Involucre 6-8 mm, broadly campanulate; both surfaces of leaves arachnoid tomentose or adaxially glabrous |
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(26) |
+ |
Involucre 4-5 mm, narrowly campanulate or campanulate, very rarely up to 6 mm in male plants; leaves tomentose or adaxially glandular or squarrose tomentose |
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(27) |
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26 (25) |
Phyllaries dark yellow or adaxially wax-yellow; leaves spatulate or oblong-spatulate, margin flat, both surfaces tomentose. |
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34 A. yunnanensis |
+ |
Phyllaries white or pale reddish white; leaves obovate or ovate-oblong, adaxially arachnoid tomentose or glabrous, abaxially tomentose. |
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35 A. muliensis |
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27 (25) |
Leaves spatulate-obovate; phyllaries white or pale red. |
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36 A. suffruticosa |
+ |
Leaves narrower, not spatulate-obovate; phyllaries white |
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(28) |
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28 (27) |
Leaves narrowly oblong or linear-oblong, both surfaces arachnoid tomentose or abaxially densely white or yellowish white tomentose. |
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37 A. stenocephala |
+ |
Leaves linear or linear-lanceolate, adaxially arachnoid or capitate glandular or squarrose tomentose, abaxially densely gray tomentose. |
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38 A. gracilis |
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29 (24) |
Phyllary claw with glandular dots at apex; both surfaces of leaves gray or yellowish white tomentose, or only with capitate-stalked glandular hairs. |
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12 A. latialata |
+ |
Phyllary claw without glandular dots at apex |
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(30) |
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30 (29) |
Stem slender, 1-2 mm in diam., densely many clustered; leaves narrowly linear, margin undulate recurved. |
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29 A. tibetica |
+ |
Stems slender or thickish, not densely many clustered; leaf margin flat or slightly reflexed or recurved and blade not narrowly linear |
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(31) |
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31 (30) |
Leaves adaxially at first arachnoid, later glabrous, without glandular hairs, or with appressed thick hairs, abaxially usually white tomentose |
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(32) |
+ |
Leaves adaxially with lanate or squarrose or capitate-stalked glandular hairs, not glabrous |
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(34) |
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32 (31) |
Involucre 4-5 mm, campanulate; leaves 3-veined. |
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27 A. corymbifera |
+ |
Involucre 6-8 mm, broadly or narrowly campanulate; leaves 1- or 3-veined |
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(33) |
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33 (32) |
Middle leaves oblong-lanceolate, 2-3 × 0.4-1 cm, margin usually reflexed or slightly recurved, obviously 3-veined. |
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15 A. likiangensis |
+ |
Leaves oblanceolate or narrowly winged, 0.4-1.3 × 0.2-0.25 cm, margin flat, 1-veined. |
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16 A. tenuissima |
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34 (31) |
Phyllaries scarious at upper parts or apex, white, opaque, middle and lower parts membranous, somewhat transparent, apex obtuse or slightly acuminate |
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(35) |
+ |
Phyllaries all scarious, opaque |
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(36) |
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35 (34) |
Leaves oblanceolate or linear-oblong, 2.5-4.5 × 0.5-2 cm; involucre 6-7 mm. |
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14 A. hymenolepis |
+ |
Leaves linear-spatulate, 2.5-5 × 0.2-0.5 cm; involucre 4-5 mm. |
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26 A. flaccida |
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36 (34) |
Stems 50-100 cm, lower parts woody, persistent, usually with developing axillary buds or branches. |
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30 A. surculosa |
+ |
Stems 3-50 cm, herbaceous, unbranched, rarely lower parts slightly woody, with developing axillary buds or branches |
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(37) |
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37 (36) |
Both surfaces of leaves tangled to cottony-papery tomentose; phyllaries wax-yellow when dry. |
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28 A. pachylaena |
+ |
Both surfaces of leaves or abaxially white, gray, or yellowish white tomentose or arachnoid, or adaxially with squarrose or capitate-stalked glandular hairs |
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(38) |
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38 (37) |
Plants with thickish woody roots or rhizome; stems unbranched |
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(39) |
+ |
Plants with long slender rhizome |
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(46) |
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39 (38) |
Both surfaces of leaves white, gray, or cinereous tomentose, without obvious capitate glandular hairs |
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(40) |
+ |
Both surfaces of leaves with obvious capitate glandular hairs or arachnoid, or margin and abaxially or rosette leaves tomentose |
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(43) |
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40 (39) |
Involucre ca. 5 mm, phyllaries white, apex obtuse or subrounded. |
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43 A. szechuanensis |
+ |
Involucre 6-8 mm, rarely only ca. 5 mm, phyllaries white, reddish, or red-purple |
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(41) |
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41 (40) |
Phyllaries purple-red or white, usually yellowish white when dry, inner ones with apex acute; leaves cinereous tomentose; flowering and sterile stems congested to pulvinate. |
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42 A. rhododactyla |
+ |
Phyllaries milky white or somewhat red, apex obtuse or rounded; leaves densely white or gray tomentose; flowering stems and rosette leaves usually caespitose |
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(42) |
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42 (41) |
Involucre (5-)6(-7) × 5-7 mm; capitula numerous, densely compound corymbiform. |
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39 A. lactea |
+ |
Involucre ca. 7 × 8 mm; capitula 4-8, densely corymbiform or headlike. |
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40 A. batangensis |
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43 (39) |
Leaves (dried) thin, yellowish green, cauline leaves with glandular hairs, only arachnoid along veins; phyllaries yellowish white, apex obtuse or rounded. |
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18 A. virens |
+ |
Leaves thick, both surfaces with arachnoid and glandular hairs; phyllaries white or sometimes pale red |
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(44) |
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44 (43) |
Stems 3-7 cm, rarely taller; flowering stems and rosette leaves somewhat congested to pulvinate. |
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44 A. xylorhiza |
+ |
Stems 15-50 cm, rarely shorter; flowering stems and rosette leaves somewhat clustered |
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(45) |
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45 (44) |
Lower stems woody, persistent; middle leaves elliptic or oblong-lanceolate, margin and midvein abaxially arachnoid tomentose; involucre 6-7 mm, phyllaries ivory white or somewhat reddish. |
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19 A. elegans |
+ |
Stems herbaceous; middle leaves oblanceolate-oblong or linear, both surfaces arachnoid tomentose; involucre 5-7 mm, phyllaries white. |
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41 A. souliei |
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46 (38) |
Lower stem slightly woody, with tomentose axillary buds or short branches; leaves adaxially arachnoid, later glabrous, abaxially thinly arachnoid tomentose and glandular, upper leaves usually tomentose on both surfaces. |
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17 A. deserti |
+ |
Stems unbranched, or sometimes branched from broken stems; leaves adaxially tomentose, or with squarrose or glandular hairs |
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(47) |
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47 (46) |
Involucre 6-8 mm; both surfaces of leaves gray, yellowish white, or rarely yellowish brown tomentose or lanate |
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(48) |
+ |
Involucre 4-5 mm or to 6 mm; both surfaces of leaves or abaxially white or gray tomentose |
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(50) |
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48 (47) |
Both surfaces of leaves gray tomentose, or upper stems and leaves fulvous tomentose. |
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13 A. bicolor |
+ |
Both surfaces of leaves yellowish white lanate or arachnoid lanate, 3-veined |
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(49) |
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49 (48) |
Leaves lingulate or linear-oblong, gradually smaller upward. |
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20 A. chungtienensis |
+ |
Leaves oblanceolate, gradually larger upward. |
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31 A. larium |
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50 (47) |
Both surfaces of leaves dark cinereous, densely tomentose and with capitate glandular hairs |
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(51) |
+ |
Leaves adaxially with squarrose or capitate glandular hairs, abaxially or both surfaces white or gray tomentose |
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(52) |
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51 (50) |
Leaves obovate or oblanceolate-spatulate; capitula numerous, congested in compound corymb. |
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22 A. spodiophylla |
+ |
Leaves broadly elliptic or subrounded; capitula few, arranged in compound corymb. |
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23 A. yangii |
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52 (50) |
Involucre semispherical; stems 10-18 cm; Taiwan. |
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24 A. transnokoensis |
+ |
Involucre campanulate or subobconic; stems 20-60 cm |
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(53) |
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53 (52) |
Involucre 4-5 mm, phyllaries white or sometimes pale red; stems densely leafy, internode 0.5-2 cm; leaves 1- or 3-veined. |
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21 A. sinica |
+ |
Involucre 5-6 mm or more, phyllaries white or yellowish white; upper stems sparsely leafy, internode 4-10 cm; leaves 3- or 5-veined. |
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25 A. aureopunctata |
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