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FOC | Family List | FOC Vol. 15 | Primulaceae

7. Androsace Linnaeus, Sp. Pl. 1: 141. 1753.

点地梅属 dian di mei shu

Androsace zambalensis

Credit: Harvard University Herbaria

Herbs perennial, annual, or biennial, acaulescent, rarely caulescent with ascending or decumbent shoots from a caudex. Leaves forming a rosette, rarely alternate; rosettes solitary or clustered, forming lax mats or compact cushions. Inflorescences umbellate, rarely a solitary flower, with bracts. Flowers 5-merous, homostylous. Calyx campanulate to subglobose, shallowly to deeply lobed. Corolla white, pink, purple, or dark red, rarely yellow; tube usually ± inflated, ca. as long as to shorter than calyx; throat constricted; lobes entire or emarginate. Stamens included, inserted on corolla tube; filaments very short; anthers ovate, apex obtuse. Style not longer than corolla tube. Capsule subglobose, dehiscing nearly to base. Seeds few to many.

Approximately 100 species widely distributed in temperate regions of the northern hemisphere; 73 species native to China.


Key 4

1 Leaves (at least outer) cartilaginous on margin or apex.   (2)
+ Leaves not cartilaginous on margin or apex.   (9)
       
2 (1) Leaves homomorphic or obscurely heteromorphic.   (3)
+ Leaves distinctly dimorphic.   (5)
       
3 (2) Leaf blade of outer leaves ovate to broadly ovate, 2-3.5 mm, apex recurved   30 Androsace squarrosula
+ Leaf blade linear to broadly lanceolate, 5-50 mm, erect.   (4)
       
4 (3) Flowers 4-7(-10) in umbels; scapes 5-10 mm   31 Androsace longifolia
+ Flowers 1 or 2; scapes to 5 mm   32 Androsace alaschanica
       
5 (2) Outer leaves spiny at apex   25 Androsace spinulifera
+ Outer leaves not spiny.   (6)
       
6 (5) Flowers 2 to many, in terminal umbels.   (7)
+ Flowers solitary, rarely 2.   (8)
       
7 (6) Bracts leaflike, cartilaginous at margin   26 Androsace graminifolia
+ Bracts not leaflike, not cartilaginous at margin   27 Androsace mariae
       
8 (6) Leaf blade linear to subulate-lanceolate; corolla white   28 Androsace lehmannii
+ Leaf blade oblong to sublinear; corolla purple   29 Androsace cernuiflora
       
9 (1) Leaves dimorphic or trimorphic, inner leaves distinctly longer than outer.   (10)
+ Leaves subequal or homomorphic.   (28)
       
10 (9) Leaves trimorphic.   (11)
+ Leaves dimorphic.   (19)
       
11 (10) Plants densely cespitose; rosettes solitary or superimposed on old rosettes, without intervals.   (12)
+ Plants laxly cespitose, with many open rosettes joined by slender shoots.   (14)
       
12 (11) Pedicel ca. 2 X as long as bracts or more   35 Androsace strigillosa
+ Pedicel shorter than bracts.   (13)
       
13 (12) Inner leaves 15-35 mm, hirsute   33 Androsace adenocephala
+ Inner leaves 5-15 mm, strigillose   34 Androsace cuttingii
       
14 (11) Plants densely white sericeous   36 Androsace nortonii
+ Plants sparsely hirsute, pilose, or pubescent.   (15)
       
15 (14) Inner leaves less than 3.5 mm wide, petiole indistinct.   (16)
+ Inner leaves 4-13 mm wide, distinctly petiolate.   (17)
       
16 (16) Shoots hirsute; outer leaves ovate-lanceolate, sheathlike basal part 2-3 × as wide as upper part 40. A. rigida   (16)
+ Shoots pilose; outer leaves spatulate, sheathlike basal part only slightly wider than upper part   41 Androsace minor
       
17 (15) Inner leaves elliptic to obovate-elliptic, villous   39 Androsace limprichtii
+ Inner leaves ovate to ovate-elliptic or suborbicular, sparsely pubescent.   (18)
       
18 (17) Calyx sparsely pubescent; inner leaves ovate-elliptic to suborbicular, sparsely pubescent   37 Androsace hookeriana
+ Calyx glabrescent, except for ciliate margin; inner leaves ovate, densely pubescent   38 Androsace ovalifolia
       
19 (10) Leaves white sericeous; stolons 5-8 cm, rust-colored villous   42 Androsace sarmentosa
+ Plants tomentose, hirsute, or pubescent, without stolons, usually with shoots.   (20)
       
20 (19) Corolla pale yellow; bracts ligulate-oblong, leaflike   43 Androsace flavescens
+ Corolla pink or purple; bracts not as above.   (21)
       
21 (20) Calyx glabrous except for ciliate margin.   (22)
+ Calyx pubescent.   (23)
       
22 (21) Leaf rosette 1 or 2-4 on caudex, without sterile shoots   44 Androsace sublanata
+ Leaf rosette forming lax mats, with sterile shoots from flowering rosette   45 Androsace forrestiana
       
23 (21) Inner leaves densely hirtellous, hairs very short.   (24)
+ Inner leaves long white hirsute, at least on margin.   (25)
       
24 (23) Leaves distinctly dimorphic, outer leaves ca. 1/3 as long as inner or shorter, with long white hairs   46 Androsace wardii
+ Leaves obscurely dimorphic, outer leaves slightly smaller than inner, ciliate   47 Androsace zayulensis
       
25 (23) Inner leaves glabrous adaxially, margin ciliate, with long and very short hairs intermixed   48 Androsace euryantha
+ Inner leaves pubescent on both surfaces.   (26)
       
26 (25) Calyx pubescent along veins and near apex of lobes   27 Androsace mariae
+ Calyx pubescent throughout.   (27)
       
27 (26) Inner leaves elliptic to suborbicular; calyx lobes ovate   50 Androsace laxa
+ Inner leaves spatulate to oblanceolate; calyx lobes triangular   51 Androsace stenophylla
       
28 (9) Inner leaves with long soft white hairs, usually very dense and tufted on distal 1/2.   (29)
+ Inner leaves with coarse hairs or glabrescent, or only fimbriate-ciliate.   (37)
       
29 (28) Leaf rosettes 2-3 mm in diam., superimposed, forming columns (leaves dimorphic in A. tapete).   (30)
+ Leaf rosettes more than 3 mm in diam., not forming columns.   (31)
       
30 (29) Calyx villous outside   61 Androsace selago
+ Calyx villous only on lobe margin   62 Androsace tapete
       
31 (29) Scapes 1- or 2-flowered, shorter than pedicel; leaf rosettes 4-8 mm in diam.   63 Androsace globifera
+ Scapes with an umbel, 2- to many flowered, distinctly longer than pedicel; leaf rosettes more than 10 mm in diam.   (32)
       
32 (31) Pedicel at most slightly longer than bracts.   (33)
+ Pedicel at least 2 × as long as bracts.   (34)
       
33 (32) Leaf rosettes forming loose tufts, internodes of shoots 1-2 × as long as leaf rosettes; corolla white, becoming pink; bracts elliptic to ovate-lanceolate   64 Androsace ovczinnikovii
+ Leaf rosettes forming dense mats; internodes of shoots usually shorter than leaf rosettes; corolla cream-colored with a pale purple or yellow eye; bracts linear to lanceolate   65 Androsace incana
       
34 (32) Scapes 8-15 cm; pedicel 5-10 × as long as bracts   66 Androsace rockii
+ Scapes 0.5-5 cm; pedicel 2-3 × as long as bracts.   (35)
       
35 (34) Leaf rosettes superimposed, without intervals; midvein and margin of leaves ± raised abaxially   67 Androsace bisulca
+ Leaf rosettes not superimposed, with intervals between rosettes; midvein and margin of leaves not raised abaxially.   (36)
       
36 (35) Plants with leaves in tight globose rosettes; scapes long white sericeous   68 Androsace robusta
+ Plants with leaves in lax rosettes; scapes spreading hirtellous   69 Androsace pomeiensis
       
37 (28) Flowers solitary.   (38)
+ Flowers 2 to many, in umbels.   (41)
       
38 (37) Leaf blade broadly lanceolate to lanceolate, glabrescent   52 Androsace tanggulashanensis
+ Leaf blade liguliform to elliptic, pubescent at least on margin.   (39)
       
39 (38) Leaf rosettes superimposed without intervals, forming columns, 2.5-4 mm in diam.   53 Androsace ciliifolia
+ Leaf rosettes at intervals, not forming columns, 5-8 mm in diam.   (40)
       
40 (39) Leaf blade obovate, adaxially glabrescent, abaxially fimbriate-ciliate   54 Androsace delavayi
+ Leaf blade narrowly obovate to spatulate, inner leaves white villous on both surfaces   63 Androsace globifera
       
41 (37) Leaf rosettes more than 1.5 cm in diam.; inner leaves distinctly longer than outer   55 Androsace lehmanniana
+ Leaf rosettes less than 1.3 cm in diam., inner leaves ca. as long as outer.   (42)
       
42 (41) Inner leaves subobovate, 1.5-2.5 mm wide, fimbriate-ciliate at apex.   (43)
+ Inner leaves linear to oblong or elliptic, 0.6-1.5 mm wide, not fimbriate ciliate at apex.   (44)
       
43 (42) Inner leaves ± incurved; bracts usually 2, ca. 2 × as long as pedicel   54 Androsace delavayi
+ Inner leaves flat; bracts several, shorter than pedicel   60 Androsace mollis
       
44 (42) Leaves hirtellous.   (45)
+ Leaves glabrous at least abaxially.   (46)
       
45 (44) Leaf rosettes globose, on nodes of shoots with internodes   56 Androsace zambalensis
+ Leaf rosettes not globose, leaves overlapping on branched caudicules   57 Androsace coronata
       
46 (44) Outer leaves whitish, scarious when dry; bracts ovate to ovate-oblong, shorter than pedicel   58 Androsace brachystegia
+ Outer leaves deep reddish brown when dry, not scarious; bracts elliptic to oblong, longer than pedicel   59 Androsace yargongensis

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