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FOC | Family List | FOC Vol. 20-21 | Asteraceae

188. Carpesium Linnaeus, Sp. Pl. 2: 859. 1753.

天名精属 tian ming jing shu

Perennials or rarely annuals. Stem not winged, without resin canals. Leaves alternate, entire or toothed, sessile or petiolate; petioles often winged. Capitula disciform, heterogamous, terminal or axillary, sessile or shortly pedunculate, usually nodding, solitary or few together, or many in spiciform racemes. Involucres subglobose or depressed-globose; phyllaries 3- or 4-seriate, outer ones herbaceous or with leaflike tips, inner ones dry, broad, obtuse. Receptacle flat, glabrous. Marginal florets female, tubular to miniradiate, in 2 to several series, fertile, slender; corollas 3-5-toothed. Disk florets bisexual, fertile, slender, many; corolla limb slightly dilated, 4- or 5-toothed, yellowish. Anthers ecalcarate, with branched tails. Endothecial tissue radial. Pollen spines with a cavity. Style branches with acute sweeping hairs not reaching furcation. Achenes ellipsoid, glabrous, longer than corolla, ribbed, with a short glandular beak, crowned by a cartilaginous ring, with elongated crystals. Pappus absent. x = 10, 18.

About 20 species: Asia, Europe; 16 species (six endemic) in China.

1 Outer phyllaries membranous or subherbaceous, shorter than inner ones, not similar to bracteal leaves   (2)
+ Outer phyllaries herbaceous or leafy, equal to or longer than inner ones, similar to bracteal leaves   (8)
2 (1) Corolla sparsely pubescent; capitula 6-10 mm in diam., terminal on stems or branches, with distinct peduncles.   1 C. divaricatum
+ Corolla glabrous; capitula 3-5 mm in diam., axillary, sessile or shortly pedunculate   (3)
3 (2) Capitula 3-6 mm in diam., with distinct peduncles or subsessile   (4)
+ Capitula 6-8 mm in diam., sessile or subsessile   (6)
4 (3) Leaves long decurrent to winged petiole.   2 C. tracheliifolium
+ Leaves shortly cuneate at base, abruptly narrowed to wingless petiole   (5)
5 (4) Stems 10-30 cm; leaves elliptic or elliptic-lanceolate, usually glabrous and dotted with glands; peduncles distinct.   3 C. minus
+ Stems 30-80 cm; leaves ovate to ovate-lanceolate, both surfaces pubescent; axillary capitula subsessile.   4 C. faberi
6 (3) Lower cauline leaves broadly ovate, base cordate or rounded.   5 C. szechuanense
+ Lower cauline leaves elliptic to lanceolate, base attenuate   (7)
7 (6) Lower cauline leaves broadly elliptic to oblong, densely pubescent.   6 C. abrotanoides
+ Lower cauline leaves oblong-lanceolate to lanceolate, glabrous.   7 C. longifolium
8 (1) Capitula campanulate, 4-10 mm in diam.   8 C. triste
+ Capitula cupuliform, over 10 mm in diam   (9)
9 (8) Corolla glabrous; plants sparsely pubescent; rarely corolla sparsely pubescent and plants densely lanate   (10)
+ Corolla pubescent; plants pubescent or densely yellowish pilose   (12)
10 (9) Capitula 2.5-3.5 cm in diam.   9 C. macrocephalum
+ Capitula 1-2 cm in diam   (11)
11 (10) Phyllary apex obtuse; lower leaves decurrent to petiole at base.   10 C. cernuum
+ Phyllary apex acute; lower leaves rounded, truncate, or cordate at base, not decurrent to petiole.   11 C. nepalense
12 (9) Lower cauline leaves broadly ovate, base cordate or rounded; petiole wingless.   12 C. cordatum
+ Lower cauline leaves elliptic, base decurrent to petiole   (13)
13 (12) Cauline leaves semiamplexicaul; plants tomentose-pilose; corolla sparsely pubescent.   13 C. velutinum
+ Cauline leaves petiolate or sessile, base not amplexicaul; corolla densely pubescent   (14)
14 (13) Bracteal leaves and outer phyllaries spatulate or linear-spatulate, apex rounded or obtuse, densely pubescent.   14 C. scapiforme
+ Bracteal leaves and outer phyllaries lanceolate, apex acuminate, sparsely pubescent   (15)
15 (14) Stems 12-35 cm tall, tomentose-pilose; capitula shortly pedunculate or subsessile; phyllaries usually not reflexed.   15 C. humile
+ Stems 35-70 cm tall, sparsely pubescent; capitula long pedunculate; phyllaries reflexed.   16 C. lipskyi

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