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FOC | Family List | FOC Vol. 14 | Clethraceae

1. Clethra Linnaeus, Sp. Pl. 1: 396. 1753.

桤叶树属 qi ye shu shu

Clethra delavayi

Credit: Harvard University Herbaria

Morphological characters and geographic distribution are the same as for the family.

Clethra sleumeriana K. S. Hao (Repert Spec. Nov. Regni Veg. 42: 84. 1937) was placed in the synonymy of C. fargesii by Hu (J. Arnold Arbor. 41: 164–190. 1960) but was resurrected by Sleumer (Bot. Jahrb. Syst. 87: 36–116. 1967). The known type material (Hunan: “Tse ching shan, in der Nähe von Sü pu,” 1000 m, fl. 8 Sep 1929, S. S. Sin 615, holotype, B) was destroyed during World War II. Hu did not see type material of this species, and Sleumer, who apparently did see it before it was destroyed, states that there are several characters that distinguish C. sleumeriana from C. fargesii. Based on Sleumer’s description, we agree that the species is likely not C. fargesii; we cannot, however, clearly delimit it from a multi-racemed form of C. delavayi, a species that, like C. sleumeriana, occurs in Hunan Province.

1 Leaf blade thickly papery to subleathery, margin of larger leaves with 0–19(–31 in C. fabri) teeth per side; plants evergreen (C. fabri apparently deciduous in SE portion of range).   (2)
+ Leaf blade papery, margin of larger leaves with 22–60 or more teeth per side; plants deciduous.   (4)
2 (1) Rachis of racemes and pedicels strigose with simple hairs; racemes 1 per branchlet; filaments hirsute; anthers 2.1–2.8 mm; style 5–7 mm at anthesis.   1 Clethra bodinieri
+ Rachis of racemes and pedicels stellate-hirsute; racemes (1 or)2–7 per branchlet; filaments glabrous; anthers 1.0–1.8 mm; style 1.5–5 mm at anthesis.   (3)
3 (2) Leaf blade abaxially often pubescent along midvein and in axils of secondary veins, otherwise glabrous; sepals suborbicular to ovate, 2.5–3 × 1.5–1.8 mm, midvein abaxially indistinct at anthesis; stamens slightly exserted or ± equal in length to petals.   2 Clethra fabri
+ Leaf blade abaxially densely grayish white appressed stellate-tomentose throughout; sepals ovate-deltoid, 1.5–2.5 × 0.8–1.2 mm, midvein usually abaxially distinct or sepals keeled at anthesis; stamens included.   3 Clethra petelotii
4 (1) Racemes 1(–4) per branchlet; petal margins neither thickened nor apically distinctly fimbriate; persistent style 7–11 mm; petals (4–)5–12 mm.   4 Clethra delavayi
+ Racemes (1–)3–8(–11) per branchlet; petal margins thickened and apically fimbriate; persistent style 3–6 mm; petals 3–5 mm.   (5)
5 (4) Racemes stellate-hirsute or stellate-subvillous; thickest rachis on each branchlet basally 1.6–2.8 mm in diam.   5 Clethra kaipoensis
+ Racemes stellate-tomentose but sometimes sparsely so; thickest rachis on each branchlet basally 0.8–1.7(–1.8) mm in diam.   (6)
6 (5) Leaf blade secondary veins (8–)11–14 on each side of midvein; sepals ovate, outside surface (visible through pubescence) usually medially blackened, midvein usually indistinct at anthesis, apex obtuse; filaments glabrous or occasionally sparsely pilose; ovary and capsule apically hirsute with longer hairs 0.7–1.1 mm.   6 Clethra barbinervis
+ Leaf blade secondary veins (11–)15–21 on each side of midvein; sepals ovate-deltoid to rarely ovate, outside surface (when visible) not medially blackened, midvein usually distinct at anthesis, apex acuminate-mucronate to rarely obtuse-mucronate; filaments basally pilose or occasionally glabrous; ovary and capsule apically hirsutulous with longer hairs 0.2–0.7 mm.   7 Clethra fargesii

Lower Taxa


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