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38. Gymnadenia R. Brown in W. T. Aiton, Hortus Kew., ed. 2. 5: 191. 1813.

手参属 shou shen shu

Authors: Xinqi Chen, Stephan W. Gale & Phillip J. Cribb

Nigritella Richard.

Herbs, terrestrial, medium-sized, slender to robust. Tubers palmately lobed, fleshy, neck with several slender, slightly fleshy roots. Stem erect, terete, fleshy, glabrous, leafy. Leaves cauline, alternate, linear-ligulate to elliptic, basally conduplicate and sheathing stem. Inflorescence capitate to elongate, with a few lanceolate sterile bracts proximally and many flowers in a dense terminal raceme; floral bracts shorter than, equal to, or slightly longer than ovary. Flowers resupinate, small to medium-sized; ovary usually twisted, narrowly cylindric or fusiform, glabrous. Sepals free, spreading; dorsal sepal concave; lateral sepals reflexed. Petals connivent with dorsal sepal and forming a hood, straight, slightly shorter than sepals; lip broadly rhombic to obovate, concave at base, spurred, conspicuously 3-lobed to entire; spur often slightly arcuate, cylindric, longer or shorter than ovary, apex obtuse or bilobed. Column short; anther oblong or ovoid, 2-locular; pollinia 2, clavate, granular-farinaceous, sectile, each with a slender caudicle attached to a separate viscidium; viscidia naked, narrowly oblong to elliptic; rostellum small to elongate, inconspicuously 3-lobed; staminodes 2, small, subglobose, placed laterally at base of anther; stigma distinctly 2-lobed, rather large, placed at base of column. Capsule erect.

About 16 species: throughout Europe and parts of C and E Asia, the Himalayas, and Japan; five species (three endemic) in China.

1 Spur 7-14 mm, ca. as long as or exceeding ovary   (2)
+ Spur 2-5 mm, much shorter than ovary   (3)
2 (1) Leaves linear-lanceolate, narrowly oblong, or lorate, 0.8-2(-2.5) cm wide; mid-lobe of lip usually smaller than lateral lobes.   1 G. conopsea
+ Leaves elliptic to elliptic-oblong, 2.5-4.5 cm wide; mid-lobe of lip equal to or slightly larger than lateral lobes.   2 G. orchidis
3 (1) Plants 50-70 cm tall; flowers yellowish green; spur shallowly 2-lobed at apex.   5 G. bicornis
+ Plants 7-50 cm tall; flowers white or pink; spur undivided, apex obtuse   (4)
4 (3) Flowers pink, rarely tinged with white; lip broadly obovate, 3-lobed at apex.   3 G. crassinervis
+ Flowers white; lip rhombic-ovate, obscurely 3-lobed at apex.   4 G. emeiensis

Lower Taxa


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