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63. Indigofera Linnaeus, Sp. Pl. 2: 751. 1753.

木蓝属 mu lan shu

Authors: Xinfen Gao & Brian D. Schrire

Shrubs, shrublets, perennial herbs, or rarely annual herbs or small trees; trichomes typically medifixed (T-shaped), equally to very unequally 2-branched and sometimes crisped, or rarely simple multicellular hairs present with glandular tips. Stipules persistent or caducous. Leaves usually imparipinnate but for some species simple or reduced to 1 leaflet; stipels present or absent; leaflet blades usually opposite but sometimes subopposite or alternate, rarely with glands, margin entire. Racemes axillary; bracts usually cadu­cous. Calyx campanulate or cup-shaped; teeth 5, subequal or abaxial one longer. Corolla usually reddish, sometimes white or yellow; standard usually covered outside with appressed trichomes but occasionally glabrous, base shortly clawed, apex usually obtuse to emarginate and mucronate; wings narrow, base auricled; keel falcate or spatulate, with spur adnate to wings. Stamens 10, diadel­phous, only vexillary one free; anthers uniform, basifixed or subbasifixed, sometimes both ends hairy, apex apiculate; pollen 3-col­porate. Ovary sessile, with 1 to many ovules; style linear, usually glabrous; stigma capitate. Legume linear, oblong, or ovoid, cylin­dric or 4-sided in cross section, 2-valved, septate, dehiscent; endocarp often spotted with tannin deposits. Seeds globose to rectan­gular; hilum small; cotyledons unequal; radicle oblique.

About 750 species: tropics and subtropics worldwide; 79 species (45 endemic, two introduced) in China.

A single collection has been seen from Yunnan of what appears to be an additional species closely related to Indigofera laxiflora Craib (Kew Bull. 1912: 148. 1912), otherwise only known from Thailand.


1 Leaves simple or 1-foliolate   (2)
+ Leaves 3- or more foliolate   (7)
       
2 (1) Leaf blade abaxially with disk-shaped or dotlike glands   (3)
+ Leaf or leaflet blade abaxially without any glands or pearl bodies   (4)
       
3 (2) Main root not tuberous; leaf blade ovate-elliptic to obovate-orbicular, abaxially with small dotlike pearl bodies, medifixed trichomes perpendicular to margin on both surfaces; legume falcate, adaxial suture echinate.   1 I. nummulariifolia
+ Main root tuberous; leaf blade narrowly obovate-lanceolate, abaxially with sparse disk-shaped glands, medifixed trichomes ± parallel to margin on both surfaces; legume straight, sutures narrowly winged, not echinate.   2 I. squalida
       
4 (2) Leaf or leaflet blade 0.4-3 × 0.3-0.6 cm; legume globose to shortly oblong, 1- or 2(or 3)-seeded   (5)
+ Leaf or leaflet blade 3-10 × 2-5.5 cm; legume cylindric, more than 3-seeded   (6)
       
5 (4) Leaf or leaflet blade usually linear but sometimes elliptic to ovate-elliptic, adaxially with appressed medifixed trichomes; legume 1-seeded.   5 I. linifolia
+ Leaf blade oblong to cordate, adaxially with long spreading white subbasifixed trichomes; legume (1 or)2-seeded.   6 I. cordifolia
       
6 (4) Leaf or leaflet blade abaxially with very unequally 2-branched trichomes; corolla standard outside with stiff brown 2-branched trichomes; stamens 8-10 mm.   7 I. caloneura
+ Leaflet blade abaxially with medifixed trichomes; corolla standard outside with soft white 2-branched trichomes; stamens 6-7 mm.   8 I. shipingensis
       
7 (1) Leaves 3-foliolate, leaflet blades abaxially with red to brown disk-shaped glands; legume sutures usually narrowly winged   (8)
+ Leaves more than 3-foliolate, or if 3-foliolate then leaflet blades without glands; legume sutures not narrowly winged   (9)
       
8 (7) Branches trailing; inflorescence longer than leaf; pedicel 3-5 mm.   3 I. pedicellata
+ Branches erect to ascending; inflorescence shorter than leaf; pedicel 0.5-1 mm.   4 I. trifoliata
       
9 (7) All leaflets alternate   (10)
+ Leaflets opposite or subopposite   (13)
       
10 (9) Racemes 0.5-3 cm; peduncle to 8 mm or absent   (11)
+ Racemes 1.8-16 cm; peduncle 9-18 mm   (12)
       
11 (10) Leaflet blades obovate to narrowly obovate; legume oblong, 0.3-0.8 cm, 1-3-seeded.   9 I. linnaei
+ Leaflet blades linear to sometimes narrowly oblong; legume linear, 1.5-2.5 cm, 7-10-seeded.   11 I. litoralis
       
12 (10) Leaves 5-9-foliolate; stamens 4-5 mm.   10 I. hendecaphylla
+ Leaves 4-6-foliolate; stamens 3-3.5 mm.   12 I. taiwaniana
       
13 (9) Stems, pedicels, and calyces with gland-tipped trichomes   (14)
+ Plants without gland-tipped trichomes   (16)
       
14 (13) Leaflet blades 0.4-1.4(-1.8) cm; racemes 2-6(-8) cm; stamens 3-4 mm.   13 I. colutea
+ Leaflet blades 1.5-3 cm; racemes 12-18 cm; stamens 5.5-7 mm   (15)
       
15 (14) Calyx 3-4.5 mm, softly pubescent, lobes more than 2 × as long as tube and filiform; leaflet blades without gland-tipped trichomes; petioles and pedicels pilose.   14 I. dolichochaete
+ Calyx 2-2.5(-3) mm, strigose, lobes up to 2 × length of tube and linear-lanceolate; leaflet blades with gland-tipped trichomes along margin and abaxial midvein; petioles and pedicels tomentose.   15 I. scabrida
       
16 (13) Stems, leaf rachises, and peduncles with ± spreading, basifixed to medifixed trichomes or crisped multicellular trichomes   (17)
+ Stems, leaf rachises, and peduncles glabrous or with appressed or appressed and spreading medifixed trichomes   (31)
       
17 (16) Stems, leaf rachises, and leaflet blade abaxial surfaces with spreading flattened and crinkly multicellular trichomes; stamens 1.2-1.3 cm.   26 I. parkesii
+ Stems, leaf rachises, and peduncles with spreading basifixed to medifixed 2-branched trichomes; stamens either less than 1.2 cm or greater than 1.3 cm (but may fall within 1.2-1.3 cm for I. dumetorum, I. esquirolii, and I. penduloides)   (18)
       
18 (17) Corolla standard 4-5 mm; calyx teeth 3-5 × as long as tube; corolla ± as long as calyx; stamens 3-4.5 mm.   37 I. hirsuta
+ Corolla standard 6 mm or more; calyx teeth less than 3 × as long as tube; corolla distinctly exserted from calyx; stamens (5.5-)6 mm or longer   (19)
       
19 (18) Racemes 2-6 cm   (20)
+ Racemes (5-)6-45 cm (5-9 cm in I. cylindracea, 5-13 cm in I. megaphylla, and 5-15 cm in I. dosua)   (22)
       
20 (19) Stamens 1.4-1.5 cm; flowers usually opening before leaves; leaves 11-19(-23)-foliolate.   39 I. pampaniniana
+ Stamens 1 cm or less; flowers opening after leaves; leaves 3-13(or 15)-foliolate   (21)
       
21 (20) Dwarf shrubs or shrublets; stems slender, with dense spreading long stiff brown trichomes and submedifixed 2-branched trichomes; stamens 9-10 mm.   38 I. argutidens
+ Shrubs; stems robust, with spreading or subspreading mixed appressed pale brown or white 2-branched trichomes; stamens 5.5-7 mm.   53 I. balfouriana
       
22 (19) Leaflet blades 0.7-2 cm   (23)
+ Leaflet blades 2-8 cm   (26)
       
23 (22) Shrubs 1-3 m tall; pedicel ca. 1.5 mm; stamens 9-10.5 mm.   42 I. stachyodes
+ Shrubs 1 m tall or less; pedicel 3-6 mm; stamens 6.5-8 mm (sometimes to 9 mm in I. dosua)   (24)
       
24 (23) Ovary and legume hairy; leaves ± sessile.   43 I. dosua
+ Ovary and legume glabrous; leaves with a 5-10 mm petiole   (25)
       
25 (24) Stipules caducous, 2-4 mm; leaves (9 or)11-15-foliolate; racemes 5-9 cm.   46 I. cylindracea
+ Stipules persistent, 5-10 mm; leaves 17-25-foliolate; racemes ca. 16 cm.   47 I. lacei
       
26 (22) Corolla purple or pink (not stated for I. penduloides)   (27)
+ Corolla white or yellowish   (29)
       
27 (26) Stamens 0.5-0.6 cm; peduncle 0.3-0.5 cm; pedicel 0.5-1 mm; ovary densely pubescent; leaves (1 or)3- or 5(or 7)-foliolate.   52 I. megaphylla
+ Stamens 1.1-1.4 cm; peduncle 3.5-6 cm; pedicel 2-3 mm; ovary glabrous   (28)
       
28 (27) Leaves 17-23-foliolate; stamens 1.1-1.2 cm.   44 I. penduloides
+ Leaves 5-9-foliolate; stamens 1.25-1.4 cm.   45 I. jikongensis
       
29 (26) Leaflet blades adaxially glabrous; calyx 2-3 mm; stamens 0.6-0.8 cm.   36 I. caudata
+ Leaflet blades adaxially hairy; calyx 4.5-6 mm; stamens 1.2-1.6 cm   (30)
       
30 (29) Leaves 9-15-foliolate; leaflet blade apices rounded to truncate; peduncle 1-1.3 cm; legume ca. 4.5 cm.   40 I. esquirolii
+ Leaves 5-9-foliolate; leaflet blade apices obtuse to acute; peduncle 2-3 cm; legume ca. 7 cm.   41 I. dumetorum
       
31 (16) Leaves 3- or 5-foliolate (5-9-foliolate in I. balfouriana and I. muliensis)   (32)
+ Leaves 5-50-foliolate (5- or 7-foliolate in I. byobiensis, 5-9-foliolate in I. muliensis, I. calcicola, and I. lenticellata)   (36)
       
32 (31) Stems, petioles, and peduncles glabrous; stamens 5-6.5 mm.   51 I. muliensis
+ Stems, petioles, and peduncles with 2-branched trichomes; stamens 6-9 mm   (33)
       
33 (32) Leaves ca. 2 cm, 3-foliolate; pedicel ca. 0.8 mm; stamens 6.5-7.5 mm.   79 I. neosericopetala
+ Leaves 3-9 cm, 3-9-foliolate; pedicel 1-3 mm   (34)
       
34 (33) Stems, petioles, and peduncles with both appressed and spreading 2-branched trichomes; stamens 6-7 mm.   53 I. balfouriana
+ Stems, petioles, and peduncles with only appressed 2-branched trichomes; stamens 7-9 mm   (35)
       
35 (34) Subshrubs; leaflets with appressed trichomes; corolla purplish red to purple, standard 6.5-8 mm; stamens 7-8 mm; ovary glabrous.   54 I. sticta
+ Shrubs; leaflets with spreading 2-branched trichomes especially dense and white; corolla pinkish, standard 8.5-11 mm; stamens 8-9 mm; ovary densely appressed white hispid.   55 I. dichroa
       
36 (31) Ovary and legume glabrous (legume glabrescent in I. hamiltonii and I. atropurpurea)   (37)
+ Ovary and legume hairy   (48)
       
37 (36) Corolla standard outside glabrous or with trichomes only at margin (glabrescent in I. hamiltonii)   (38)
+ Corolla standard outside hairy   (41)
       
38 (37) Bracts to 3.5 mm wide, boat-shaped; racemes sessile or subsessile, with persistent bud scales at base.   23 I. cassioides
+ Bracts 1-1.5 mm wide, ovate, ovate-lanceolate, or linear-lanceolate; racemes pedunculate, without persistent bud scales at base   (39)
       
39 (38) Stamens 1.2-1.4 cm; pedicel 3-5 mm.   16 I. kirilowii
+ Stamens 0.5-0.8 cm; pedicel 1-2 mm   (40)
       
40 (39) Leaves 3- or 5(-11)-foliolate, to 2.2 × 1.2 cm; racemes much longer than leaves.   24 I. hamiltonii
+ Leaves (7-)11-19(or 21)-foliolate, to 6.5 × 2.5 cm; racemes equal to slightly longer than leaves.   25 I. atropurpurea
       
41 (37) Bracts 4-6 × 2-3 mm, cuspidate   (42)
+ Bracts 1-5 × 1-1.5 mm, linear-lanceolate to ovate   (43)
       
42 (41) Subshrubs to 0.9 m; stems and branches mostly glabrous; leaflet blades 1.2-2.5(-3.2) × 0.6-1 cm, adaxially hairy; stamens 7-8 mm.   21 I. bracteata
+ Robust shrubs 1-3 m; stems and branches with appressed medifixed trichomes; leaflet blades 1.2-6.5(-8) × 0.7-3.5(-4.5) cm, adaxially glabrous; stamens 8-10 mm.   22 I. hebepetala
       
43 (41) Leaves (13 or)15-19-foliolate; leaflet blades 0.8-1.3 cm wide; pedicel 1-1.5 mm.   60 I. delavayi
+ Leaves 5-15-foliolate (if to 23 leaflets in I. decora, then pedicels 3-6 mm); leaflet blades to 3 cm wide; pedicel 2-6 mm   (44)
       
44 (43) Leaflet blades with appressed medifixed trichomes on both surfaces; stamens (11-)12-15 mm   (45)
+ Leaflet blades adaxially glabrous   (46)
       
45 (44) Leaflet blades usually ovate-lanceolate, ovate-oblong, or lanceolate, apex acuminate, acute, or rarely obtuse; calyx 2.5-3.5 mm; stipels ca. 1.5 mm.   18 I. decora
+ Leaflet blades elliptic to ovate-elliptic, apex obtuse; calyx 4-4.5 mm; stipels 2-4 mm.   20 I. carlesii
       
46 (44) Stamens 0.7-0.9 cm.   17 I. fortunei
+ Stamens 1.1-1.4 cm   (47)
       
47 (46) Leaflet blades ovate-lanceolate, ovate-oblong, or lanceolate, 2-7.5(-10) cm; bracts linear-lanceolate, ca. 3 mm; pedicel 3-6 mm.   18 I. decora
+ Leaflet blades ovate, ovate-rhombic, or suborbicular, 1.3-2.5 cm; bracts ovate-lanceolate, 2-2.5 mm; pedicel ca. 2.5 mm.   19 I. venulosa
       
48 (36) Leaflet blades adaxially ± glabrous (adaxially with sparse trichomes in I. bungeana, adaxially glabrous or subglabrous in I. delavayi, adaxially glabrous except for isolated appressed hairs along midrib in I. sensitiva)   (49)
+ Leaflet blades adaxially with appressed 2-branched trichomes   (54)
       
49 (48) Stamens (1.3-)1.4-1.7 cm; legume 4-5(-6) cm.   60 I. delavayi
+ Stamens 0.3-0.8 cm; legume 2-3.5 cm   (50)
       
50 (49) Stems, petioles, and peduncles glabrous.   51 I. muliensis
+ Stems, petioles, and peduncles sparsely hairy   (51)
       
51 (50) Leaves 3-9-foliolate   (52)
+ Leaves 11-31-foliolate   (53)
       
52 (51) Shrubs to 1 m tall; leaves 2.5-5 cm; leaflet blades 5-15(-25) × 3-10(-15) mm; peduncle 1-5 mm.   57 I. bungeana
+ Subshrubs to 30 cm tall; leaves 0.6-1.5(-2) cm; leaflet blades 3.5-6 × 1.5-2.5 mm; peduncle 9-15 mm.   64 I. chaetodonta
       
53 (51) Leaves 21-31-foliolate; stamens 3-4 mm.   63 I. sensitiva
+ Leaves 11-21-foliolate; stamens 5-5.5 mm.   65 I. mengtzeana
       
54 (48) Corolla standard outside with appressed stiff brown trichomes   (55)
+ Corolla standard outside with white 2-branched trichomes (unknown for I. jindongensis)   (61)
       
55 (54) Stamens 13-16 mm.   33 I. sootepensis
+ Stamens 3-7 mm   (56)
       
56 (55) Corolla standard with tuft of hairs inside at base; legume 1.7-2.8 cm.   31 I. nigrescens
+ Corolla standard without tuft of hairs inside; legume 2-9 cm   (57)
       
57 (56) Legume deflexed; stamens 3-5 mm   (58)
+ Legume spreading to erect; stamens 3-7 mm   (59)
       
58 (57) Legume 10-15 × 2.5-3 mm, falcate; stamens 3-4 mm.   27 I. suffruticosa
+ Legume 25-30 × 1-2(-2.2) mm, straight to semicircular but never falcate; stamens 4-5 mm.   30 I. tinctoria
       
59 (57) Leaflet blades ovate-lanceolate, 3-6 cm; legume spreading, 2.5-4.5 cm; seeds discoid, stacked like a pile of coins.   32 I. zollingeriana
+ Leaflet blades oblanceolate-oblong, 2-4 cm; legume ascending to erect; seeds rectangular to quadrate, arranged typically as in genus   (60)
       
60 (59) Legume 6-9 cm; racemes 5-12 cm; stamens 5-7 mm.   34 I. galegoides
+ Legume 2-3.5 cm; racemes to 26 cm; stamens 3-4 mm.   35 I. densifructa
       
61 (54) Pedicel (2-)4-9 mm; leaf rachis adaxially flattened, slightly winged   (62)
+ Pedicel 0.5-3(-4) mm; leaf rachis not flattened or winged   (63)
       
62 (61) Calyx 2-3 mm, teeth ± equaling tube.   48 I. henryi
+ Calyx 4-5 mm, teeth to 2 × as long as tube.   49 I. myosurus
       
63 (61) Racemes usually (9-)10-25(-35) cm   (64)
+ Racemes usually shorter, (1-)2-10(-15) cm   (67)
       
64 (63) Racemes usually erect or ascending   (65)
+ Racemes usually pendulous or spreading   (66)
       
65 (64) Corolla standard incurved at apex; peduncle to 0.5 cm; stamens 5-6 mm; corolla pink.   56 I. amblyantha
+ Corolla standard not incurved at apex; peduncle 0.8-5 cm; stamens 7.5-8.5 mm; corolla dark crimson-red.   75 I. szechuensis
       
66 (64) Racemes pendulous, to 30 cm; corolla standard bluish pink, densely downy white puberulent dorsally; leaves (11 or)13-23(-27)-foliolate, leaflet blades adaxially glabrous.   61 I. pendula
+ Racemes spreading ± horizontally with tips ascending; corolla standard bright reddish pink, thinly strigillose-puberulent; leaves (7 or)9-17(-23)-foliolate, leaflet blades adaxially hairy.   62 I. howellii
       
67 (63) Shrublets, 10-30 cm tall; leaflet blade venation obviously reticulate on both surfaces; stamens 4-5 mm, much shorter than 7-9 mm tapering corolla keel.   50 I. reticulata
+ Shrubs, mostly taller than 30 cm; leaflet blade venation obscure on both surfaces; stamens only ca. 1 mm shorter than corolla keel   (68)
       
68 (67) Leaves 21-31-foliolate; stamens 3-4 mm.   63 I. sensitiva
+ Leaves usually 21-foliolate or less (if more than 21-foliolate in I. franchetii, I. hancockii, I. heterantha, and I. wightii, then stamens 5-11.5 mm); stamens usually longer than 4 mm (3-4 in I. byobiensis, I. ramulosissima, and I. silvestrii and unknown in I. jindongensis)   (69)
       
69 (68) Leaves (9-)13-25(-29)-foliolate   (70)
+ Leaves usually 5-13-foliolate (occasionally to 17-foliolate in I. mairei)   (74)
       
70 (69) Stamens 5-5.5 mm   (71)
+ Stamens 6-11.5 mm   (72)
       
71 (70) Leaflet blades with appressed 2-branched trichomes on both surfaces, midvein abaxially prominent and terminating in a tubular gland; legume erect.   59 I. wightii
+ Leaflet blades with sparse trichomes abaxially and glabrescent adaxially, midvein not as above; legume spreading.   65 I. mengtzeana
       
72 (70) Petiole 1-9 mm; stamens (6-)7-11.5 mm; young racemes in bud grayish green; racemes ascending to erect (71. I. chenii will also key out here; see note under description of this species).   70 I. heterantha
+ Petiole 8-20 mm; stamens 6-7 mm; young racemes in bud dark brown or densely appressed white hairy; racemes stiffly erect   (73)
       
73 (72) Leaves (5-)9-19-foliolate; young branches, leaves, and racemes with appressed brown 2-branched trichomes.   68 I. hancockii
+ Leaves (11-)17-27-foliolate; young branches, leaves, and racemes with dense appressed grayish white 2-branched trichomes.   69 I. franchetii
       
74 (69) Legume 3.5-6 cm (unknown in I. acutipetala and I. wilsonii)   (75)
+ Legume 1.5-3 cm (to 3.5 cm in I. mairei, unknown in I. acutipetala and I. wilsonii)   (79)
       
75 (74) Petiole 1.2-5 cm; leaflet blades 1-4(-5) cm   (76)
+ Petiole 0.2-1.1(-1.3) cm; leaflet blades 0.3-1.6 cm   (77)
       
76 (75) Stems terete; leaves to 18 cm; petiole (1.5-)2-5 cm; racemes 11-15 cm.   56 I. amblyantha
+ Stems 4-ribbed; leaves 8.5-10.5 cm; petiole 1.2-1.7 cm; racemes 4-7 cm (but only known in fruit).   78 I. jindongensis
       
77 (75) Stipules 5-6(-7) mm; pedicel 2.5-4 mm; corolla standard 11-14 mm.   74 I. wilsonii
+ Stipules 2-4 mm; pedicel 1-2 mm; corolla standard 7-9.5 mm   (78)
       
78 (77) Leaves 4-10 cm; racemes 10-19 cm; stamens 7.5-8.5 mm.   75 I. szechuensis
+ Leaves ca. 3 cm; racemes 5-9 cm; stamens 5-6.5 mm.   77 I. acutipetala
       
79 (74) Stamens 6.5-13 mm (sometimes ca. 6 mm in I. mairei, 5-6.5 mm in I. acutipetala)   (80)
+ Stamens 3-6.5 mm   (85)
       
80 (79) Corolla standard outside spreading soft pubescent   (81)
+ Corolla standard outside with dense appressed straight and stiffly medifixed 2-branched trichomes   (82)
       
81 (80) Shrubs 1-2(-3) m tall; peduncle 0.1-1 cm; corolla standard 0.7-0.9(-1) cm; stamens 6-8 mm.   72 I. mairei
+ Shrubs 0.3-0.8 m tall; peduncle 1.5-4 cm; corolla standard 1.1-1.4 cm; stamens 10-13 mm.   74 I. wilsonii
       
82 (80) Corolla standard with a distinct ca. 1 mm mucro at apex; stamens 5-6.5 mm.   77 I. acutipetala
+ Corolla standard without a mucro; stamens 7-12 mm   (83)
       
83 (82) Stamens 11-12 mm; pedicel 2-4 mm.   76 I. pseudoheterantha
+ Stamens 7-9 mm; pedicel 1-2 mm   (84)
       
84 (83) Corolla standard pink to rose; peduncle 0.1-0.5(-1) cm; stipules ca. 1 mm; leaves 1.5-2.5 cm; racemes 3-6(-8) cm.   73 I. rigioclada
+ Corolla standard dark crimson-red; peduncle 0.8-5 cm; stipules 2-4 mm; leaves 4-10 cm; racemes 10-19 cm.   75 I. szechuensis
       
85 (79) Stamens 3-4 mm   (86)
+ Stamens 5-6.5 mm (4-5 mm in I. bungeana)   (88)
       
86 (85) Stems prostrate; legume 4-angled, spreading (from Taiwan only).   28 I. byobiensis
+ Stems erect; legume cylindric   (87)
       
87 (86) Racemes not spinescent (from Taiwan only).   29 I. ramulosissima
+ Racemes spinescent.   58 I. silvestrii
       
88 (85) Leaflet blades 0.5-4(-5) × 0.3-2 cm; racemes 4-15 cm   (89)
+ Leaflet blades 0.3-0.7 × 0.2-0.5 cm; racemes 1-5 cm   (90)
       
89 (88) Corolla standard apically hooded; leaves to 18 cm; terminal leaflet blade narrowly ovate to elliptic, 2-5 cm; petiole (1.5-)2-5 cm; stamens 5-6 mm.   56 I. amblyantha
+ Corolla standard not apically hooded; leaves 2.5-5 cm; terminal leaflet blade obovate to elliptic, 0.5-1.5(-2.5) cm; petiole 0.4-1.6(-2) cm; stamens 4-5 mm.   57 I. bungeana
       
90 (88) Leaflet blade indumentum very densely silvery white strigose with appressed 2-branched trichomes, obscuring surfaces.   66 I. calcicola
+ Leaflet blade indumentum less dense, abaxially not obscuring surfaces.   67 I. lenticellata

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