| 1 |
Gynophore free end (vertical part of gynobase) rising above nutlets as thick tetrahedral mucro with short style at apex. |
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(2) |
| + |
Gynophore entirely hidden between nutlets (gynophore should not be confused with the thin style sometimes elevated above nutlets as in L.brachycentra and L.microcarpa). |
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(4) |
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| 2 (1) |
Nutlets 3-5 mm, lustrous, stonelike, tuberculate, with thick walls and without ribs or disc or anchor-like tip on tubercles |
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1 Lappula spinocarpos |
| + |
Nutlets 2-3 mm, finely tuberculate, with very short glochids on margin of inconspicuous disc, without glochids on margin of well-developed disc. |
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(3) |
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| 3 (2) |
Calyx lobes exceeding fruit; pedicel always straight |
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3 Lappula occultata |
| + |
Calyx lobes short, appressed to base of much longer fruit; pedicel declining, sometimes recurved |
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2 Lappula sinaica |
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| 4 (1) |
Developed nutlets 1, the other 3 aborting |
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31 Lappula monocarpa |
| + |
Developed nutlets 4. |
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(5) |
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| 5 (4) |
Disc of nutlets narrow and nearly closed, marginal parts of nutlets involute on disc. |
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(6) |
| + |
Disc of nutlets well developed, marginal parts of nutlets not involute on disc. |
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(7) |
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| 6 (5) |
Glochids on margin of disc ca. 0.5 mm |
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4 Lappula balchaschensis |
| + |
Glochids on margin of disc ca. 1.5 mm |
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5 Lappula deserticola |
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| 7 (5) |
Disc margin of all or (1 or) 2 nutlets with well-developed mostly broad wing. |
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(8) |
| + |
Disc margin of nutlets without well-developed wing, marginal prickles of disc sometimes widened and confluent into a very narrow winglike border. |
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(16) |
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| 8 (7) |
Perennial herbs. |
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(9) |
| + |
Biennial or annual herbs (when uncertain, stems straight, tall). |
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(10) |
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| 9 (8) |
Herbs forming small tufts of leaf rosettes, with several or sometimes many lateral ascending stems; nutlets 5-7 mm across |
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33 Lappula transalaica |
| + |
Cespitose herbs, with 1 or 2 straight stems; nutlets ca. 3 mm across |
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15 Lappula caespitosa |
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| 10 (8) |
Nutlet with entire wing, pubescent or not. |
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(11) |
| + |
Nutlet with dentate wing, not pubescent. |
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(12) |
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| 11 (10) |
Nutlets tuberculate, densely pubescent, without prickles |
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32 Lappula lasiocarpa |
| + |
Nutlets neither tuberculate nor pubescent, 2 nutlets with a single row of marginal glochids |
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30 Lappula alatavica |
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| 12 (10) |
Biennial or ?perennial herbs, with well-developed lateral rosette of leaves; stems (2 or) 3-5, few branched or not, straight |
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27 Lappula ferganensis |
| + |
Annual herbs without rosette of leaves after anthesis; stems usually branched. |
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(13) |
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| 13 (12) |
All nutlets homomorphic, with narrow wings to 1 mm, sometimes 1 nutlet wingless, disc of nutlets granulose |
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19 Lappula granulata |
| + |
Nutlets heteromorphic, 2 with leathery wings and 2 with 2 rows of glochids on disc margin. |
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(14) |
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| 14 (13) |
Corolla limb (5-)5.5-6 mm |
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34 Lappula lipskyi |
| + |
Corolla limb 2.5-3.5(-4) mm. |
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(15) |
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| 15 (14) |
Marginal glochids in 2 rows, uniform in length |
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29 Lappula heteromorpha |
| + |
Marginal glochids in 1 row, some short glochids forming a second row near base of nutlets |
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28 Lappula anocarpa |
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| 16 (7) |
Style projecting 1-1.5 mm above nutlets, filiform, persistent at fruiting. |
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(17) |
| + |
Style hidden among apices of nutlets or projecting less than 0.5 mm above nutlet. |
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(18) |
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| 17 (16) |
Prickles along disc margin 0.5-0.9 mm, mostly developed in 1 row (sometimes a second row of prickles 0.1-0.2 mm, with very small tubercles and anchorlike tips) |
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10 Lappula microcarpa |
| + |
Prickles along disc margin 0.1-0.2 mm or absent and replaced by tubercles |
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11 Lappula brachycentra |
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| 18 (16) |
Disc margin of nutlets with only 1 row of glochids (sometimes a medial row of glochids present). |
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(19) |
| + |
Disc margin of nutlets with 2 or 3 rows of glochids. |
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(29) |
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| 19 (18) |
Perennial cespitose herbs, with several to many stems, branched only above |
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15 Lappula caespitosa |
| + |
Annual or rarely biennial herbs. |
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(20) |
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| 20 (19) |
Corolla sky blue, limb 5-8 mm |
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35 Lappula macrantha |
| + |
Corolla blue or white, limb to 3.5 mm. |
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(21) |
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| 21 (20) |
Corolla white; nutlets with marginal glochids to 0.5 mm, disc with 1 or 2 medial rows of glochids on tubercles |
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6 Lappula macra |
| + |
Corolla blue or white-blue; nutlets with longer marginal glochids or disc finely tuberculate. |
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(22) |
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| 22 (21) |
Heteromorphic nutlets present, 2 with short glochids, 2 with longer glochids; homomorphic nutlets sometimes also present. |
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(23) |
| + |
All nutlets homomorphic. |
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(24) |
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| 23 (22) |
All nutlets heteromorphic, disc and adaxial surface smooth |
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16 Lappula karelinii |
| + |
Nutlets in lower part of inflorescences homomorphic, disc and adaxial surface tuberculate |
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13 Lappula duplicicarpa |
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| 24 (22) |
Marginal glochids slender, 0.5-0.9 mm |
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9 Lappula tenuis |
| + |
Marginal glochids 1-2.5(-4.5) mm. |
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(25) |
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| 25 (24) |
Marginal glochids 1-2 mm; plants branched mainly on upper part of stems. |
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(26) |
| + |
Marginal glochids 2-2.5(-4.5) mm; plants usually branched basally. |
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(27) |
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| 26 (25) |
Plants 25-40(-50) cm high; marginal spines 1-1.2 mm, disc not keeled |
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8 Lappula intermedia |
| + |
Plants 15-20(-30) cm high; marginal spines ca. 1.5 mm, disc with protruding keel |
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7 Lappula stricta |
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| 27 (25) |
Marginal glochids hollow, terete |
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36 Lappula physacantha |
| + |
Marginal glochids solid, or subglobose. |
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(28) |
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| 28 (27) |
Pedicel 1.5-4 mm, usually curved; nutlets 3.5-4 mm excluding prickles |
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12 Lappula semiglabra |
| + |
Pedicel to 1 mm, straight; nutlets ca. 3 mm excluding prickles |
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14 Lappula patula |
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| 29 (18) |
Perennial herbs; stems several, 7-15 cm tall, ascending; basal leaves numerous, forming a rosette; glochids on disc 0.2-0.5 mm. |
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(30) |
| + |
Biennial or annual herbs; stems few, usually 30-45 cm tall; basal leaves few, rarely forming a rosette; glochids on disc 1-3 mm, rarely smaller. |
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(31) |
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| 30 (29) |
Herbs silvery sericeous; basal leaves linear to filiform, to 4 cm × 1-2 mm; corolla ca. 5 mm; center line of disc of nutlet without glochids |
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20 Lappula sericata |
| + |
Herbs green-gray pilose; basal leaves spatulate, to 3 cm × 2-4 mm; corolla 2-2.5 mm; center line of disc of nutlet with glochids |
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21 Lappula himalayensis |
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| 31 (29) |
Biennial herbs; corolla with spreading limb, 4-6(-8) mm across. |
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(32) |
| + |
Annual herbs; corolla with campanulate limb, 2-3.5(-4) mm across. |
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(35) |
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| 32 (31) |
About 2/3 of adaxial angle of nutlets affixed to gynobase; gynobase longer than nutlets |
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25 Lappula ramulosa |
| + |
Entire adaxial angle of nutlets affixed to gynobase; gynobase shorter than nutlets. |
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(33) |
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| 33 (32) |
Corolla sky blue, limb to 6-8 mm across, throat appendages blue or dark blue |
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22 Lappula tianschanica |
| + |
Corolla blue, limb to 4-5 mm across, throat appendages white, yellowish in herbarium specimens. |
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(34) |
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| 34 (33) |
Glochids on disc numerous but short, glochids of main row on margin of disc 1-1.5 mm; herbs to 30(-35) cm tall |
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24 Lappula pratensis |
| + |
Glochids on disc usually 3 along center line, to 1 mm, glochids of main row on margin of disc 1.5-2.5 mm; herbs to 40-45 cm tall |
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23 Lappula tadshikorum |
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| 35 (31) |
Calyx lobes stellate spreading in fruit, elongated to at least 7-8 mm; leaves light green or green, with sparse spreading hairs on large white tubercles; main marginal glochids with large anchorlike tip |
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8 Lappula intermedia |
| + |
Calyx lobes spreading in fruit, not stellate, elongated to 4-5 mm; leaves usually densely appressed pubescent, grayish green; marginal glochids with small anchorlike tip. |
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(36) |
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| 36 (35) |
Upper part of nutlets separated by deep oval or elliptic pit; style protruding above nutlets (excluding glochids) 0.5-1 mm |
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26 Lappula consanguinea |
| + |
Upper part of nutlets without pit; style protruding above nutlets to 0.2-0.3 mm. |
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(37) |
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| 37 (36) |
Main marginal prickles widened basally, confluent near base, 0.4-1.5(-2) mm |
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18 Lappula shanhsiensis |
| + |
Main marginal prickles all separated, 1.5-2 mm (or more) |
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17 Lappula myosotis |