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8. Physaliastrum Makino, Bot. Mag. (Tokyo). 28: 20. 1914.

散血丹属 san xue dan shu

Archiphysalis Kuang.

Shrubs or perennial or perhaps annual herbs; pubescence of simple hairs. Stems dichotomously branched. Leaves simple, petiolate. Inflorescences in branch forks, occasionally axillary, solitary or sometimes 2- or 3-flowered clusters; peduncle short or obsolete. Flowers mostly actinomorphic, 5-merous, long pedicellate, nodding. Calyx short campanulate or conical, lobed to halfway, lobes equal or not. Corolla broadly campanulate to rotate, lobed to halfway; lobes wide spreading, densely villous near base of tube, sometimes spotted. Nectaries sometimes present. Stamens inserted in corolla tube; filaments pubescent or glabrous; anthers dehiscing longitudinally. Disc rudimentary or absent. Ovary 2-locular; ovules numerous. Fruiting calyx becoming enlarged, applied to berry or inflated, mostly open at apex; veins obscure or thickened into angles or ribs, mostly muricate or bristly with soft emergences that appear deltate and scalelike when dry, these sometimes localized on angles. Fruit a juicy or fleshy berry enveloped in calyx. Seeds numerous, reniform-discoid, minutely pitted; embryo curved, subperipheral.

About nine species: Asia, seven species in China.

The genus Archiphysalis, which was recognized in FRPS, was placed into synonomy under Physaliastrum by D'Arcy & Zhang, Novon 2: 124-128. 1992. Many species were formerly known as Chamaesaracha A. Gray and Leucophysalis Rydberg, but these genera now include only New World taxa.


1 Veins of fruiting calyx thickened into ribs or angles, these sometimes with dentate emergences, area between veins smooth.   (2)
+ Veins of fruiting calyx obscure, evenly muricate with bristly emergences that dry deltate and scalelike.   (3)
       
2 (1) Fruiting calyx with evenly thickened ribs or angles, especially basally, lacking emergences   6 Physaliastrum sinense
+ Fruiting calyx with unevenly thickened, winglike ribs, these with teeth or emergences that dry deltate   7 Physaliastrum chamaesarachoides
       
3 (1) Corolla with an evident nectary; fruiting calyx about as long as berry, apex open   5 Physaliastrum echinatum
+ Corolla without a nectary; fruiting calyx longer than berry, apex slightly contracted.   (4)
       
4 (3) Flowering calyx 5-dentate, lobes equal, broadly deltate   1 Physaliastrum yunnanense
+ Flowering calyx 5-divided halfway, lobes unequal, narrowly deltate or oblong.   (5)
       
5 (4) Corolla 1 × 1 cm; stamens more than half as long as corolla, filaments glabrous   4 Physaliastrum sinicum
+ Corolla 1.5-2 × 1.5-2 cm; stamens half as long as corolla, filaments glabrous or pubescent.   (6)
       
6 (5) Calyx lobes subequal; filaments sparsely pubescent   2 Physaliastrum heterophyllum
+ Calyx lobes very unequal, longest 1 as long as corolla; filaments glabrous   3 Physaliastrum kweichouense

Lower Taxa


 

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