5. Physaliastrum echinatum (Yatabe) Makino, Bot. Mag. (Tokyo). 28: 21. 1914.
日本散血丹 ri ben san xue dan
Chamaesaracha echinata Yatabe, Bot. Mag. (Tokyo) 5: 355. 1891.
Plants 50-70 cm tall, sparsely pubescent or villous. Petiole 0.5-2.5 cm; leaf blade ovate to broadly ovate, 4-12 × 3-6 cm, pubescent, ciliate, base cuneate, decurrent, apex acute. Inflorescences usually 2- or 3-flowered clusters in leaf axils or branch forks. Pedicel 2-4 cm, nodding. Calyx short campanulate, 3-5 mm in diam., pubescent and villous with scalelike abaxial emergences; lobes short deltate, equal, 0.5-1.5 mm. Corolla campanulate, ca. 1 cm in diam.; lobes pubescent, ciliate, with 5 pairs of nectaries adaxially near middle part of tube. Stamens included, slightly shorter than corolla tube; filaments pubescent, ca. 6 mm; anthers 1.8-2 mm. Fruiting calyx subglobose, nearly as long as berry, open at apex, thick, muricate. Berry globose, ca. 1 cm in diam., enveloped in fruiting calyx, only apex exposed. Seeds subdiscoid. Fl. Jun-Aug, fr. Jul-Sep.
Grassy slopes, forests; low elevations. Hebei, Heilongjiang, Jilin, Liaoning, Shandong, Zhejiang [Japan, Korea, Russia]
This species was treated in FRPS as Physaliastrum japonicum (Franchet & Savatier) Honda, which is a different species.