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FOC | Family List | FOC Vol. 18 | Scrophulariaceae

35. Veronicastrum Heister ex Fabricius, Enum. 111. 1759.

腹水草属 fu shui cao shu

Botryopleuron Hemsley; CalorhabdosBentham; Leptandra Nuttall.

Herbs, perennial. Roots usually densely yellow tomentose when young. Rhizomes short to long, rarely absent. Stems erect or arching and rooting at apex. Leaves alternate, opposite, or whorled. Inflorescences terminal or axillary, spicate. Flowers usually crowded. Calyx 5-lobed, upper lobe slightly smaller than others. Corolla 4-lobed; tube tubular, straight or slightly curved, usually with a ring of villous hairs inside, rarely glabrous; limb actinomorphic or 2-lipped; lobes unequal in width, upper widest, lower narrowest. Stamens 2; filaments usually basally villous, rarely glabrous, adnate to upper part of corolla tube; anther locules connivent, not confluent. Stigma small, slightly dilated. Capsule ovoid to ovoid-globose, slightly compressed laterally, 2-grooved, 4-valved. Seeds numerous per capsule.

About 20 species: E Asia, North America; 13 species in China.

1 Stems winged; corolla tube as long as wide; corolla lobes as long as tube.   (2)
+ Stems terete or angular (but usually winged in V. brunonianum subsp. sutchuense); corolla tube longer than wide; corolla lobes shorter than tube (equal to tube in V. rhombifolia and V. axillare).   (3)
2 (1) Stems ca. 100 cm tall; leaf blade oblong, ovate, or lanceolate   10 Veronicastrum caulopterum
+ Stems 15-30 cm tall; leaf blade linear to elliptic   11 Veronicastrum formosanum
3 (1) Corolla slightly to conspicuously 2-lipped, lower lobes reflexed, upper often galeate; seed coat transparent.   (4)
+ Corolla actinomorphic, lobes straight; seed coat not transparent.   (5)
4 (3) Stems herbaceous, erect; leaves sessile, narrowly elliptic; corolla conspicuously 2-lipped, lower lobes strongly reflexed   1 Veronicastrum brunonianum
+ Stems woody, climbing; leaves petiolate, blade ovate to ovate-lanceolate; corolla slightly 2-lipped, lower lobes not strongly reflexed   2 Veronicastrum yunnanense
5 (3) Inflorescences terminal on main stems; flowers pedicellate; leaves sessile.   (6)
+ Inflorescences axillary or terminal on branches (sometimes terminal on main stem in V. stenostachium); flowers sessile; leaves short petiolate to subsessile.   (7)
6 (5) Leaves in whorls of 4-6, oblong to broadly linear, 1.5-4.5 cm wide   12 Veronicastrum sibiricum
+ Leaves alternate, linear, ca. 0.6 cm wide   13 Veronicastrum tubiflorum
7 (5) Bracts ovate-orbicular or ovate-lanceolate; corolla tube subglabrous inside   8 Veronicastrum robustum
+ Bracts linear-lanceolate to subulate; corolla tube with a ring of villous hairs inside.   (8)
8 (7) Leaves rhomboid, rarely rhomboid-ovate; corolla lobes 1/3-1/2 of corolla length   9 Veronicastrum rhombifolium
+ Leaves not rhomboid (rhomboid-ovate in V. villosulum); corolla lobes 1/6-1/4 of corolla length (1/3-1/2 in V. axillare).   (9)
9 (8) Ovaries and young fruit hairy; inflorescence axes, bracts, and calyx lobes densely capitate glandular or hispidulous   3 Veronicastrum longispicatum
+ Ovaries and young fruit glabrous; inflorescence axes, bracts, and calyx lobes not densely capitate glandular (bracts and calyx lobes sometimes densely pubescent in V. villosulum var. villosulum and var. hirsutum).   (10)
10 (9) Corolla lobes 1/3-1/2 of corolla length; stems angular in middle and apically   6 Veronicastrum axillare
+ Corolla lobes 1/6-1/4 of corolla length; stems terete, sometimes apically angular.   (11)
11 (10) Inflorescences less than 1.5 cm   7 Veronicastrum villosulum
+ Inflorescences 1.5-8 cm.   (12)
12 (11) Leaves narrowly ovate to lanceolate, 7-20 cm   4 Veronicastrum stenostachyum
+ Leaves ovate-orbicular to orbicular, 4-7 cm   5 Veronicastrum latifolium

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