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FOC | Family List | FOC Vol. 4 | Fagaceae | Lithocarpus

40. Lithocarpus corneus (Loureiro) Rehder in Bailey, Stand. Cycl. Hort. 3569. 1917.

烟斗柯 yan dou ke

Trees usually less than 15 m tall. Branchlets light tawny, dark gray, or silver-gray, glabrous or pubescent, sparsely lenticellate; lenticels raised. Leaves usually congested at apex of branches; petiole 0.5-4.5 cm; leaf blade (5-)10-15 × 2-4.5 cm, papery to leathery, concolorous, with ± translucent, minute (visible under hand lens) scalelike glands, base cuneate to subrounded and symmetric or oblique, margin dentate, shallowly undulate, or rarely entire, apex acuminate to acute; secondary veins 9-26 on each side of midvein, ending in teeth; tertiary veins abaxially slender, evident, subparallel. Male inflorescences often with female flowers borne at base of rachis; Female inflorescences less than 10 cm; cupules in clusters of ca. 3 or sometimes solitary. Infructescences 5-7; rachis 3-4 mm thick. Cupule cupular to subglobose, 2.2-4.5 × 2.5-5.5 cm, enclosing ca. 1/2 of nut, wall (1-)2-3 mm thick, woody, and basally thickened; bracts triangular to rhomboid, center and margin ridged or fused with cupule and ± united into concentric rings. Nut subglobose to turbinate, rarely glabrous, apex rounded, flat, or slightly concave, wall ± horny and usually thicker than wall of cupule; scar covering 1/2 to most of nut, convex. Cotyledons 4-8-lobed. Fl. almost all year around but mainly May-Jul, fr. maturing on 1-year-old branchlets.

Broad-leaved evergreen forests, frequent on sunny slopes and in dry places, coastal regions; below 1000 m. S Fujian, Guangdong, Guangxi, S Guizhou, Hainan, S Hunan, Taiwan, EC and SE Yunnan [NE Vietnam]

A widespread and variable species. A critical review of the pattern of variation within Lithocarpus corneus may show that the acceptance of most of the varieties below is unjustified.

1 Leaf blade abaxially hairy   40e var. hainanensis
+ Leaf blade glabrous or abaxially with short hairs only along midvein or at axils of secondary veins.   (2)
2 (1) Bracts on cupule mostly reduced and often united into a few concentric rings, slightly convex   40b var. zonatus

Lower Taxa


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