31. Cardamine engleriana O. E. Schulz, Bot. Jahrb. Syst. 32: 407. 1903.
光头山碎米荠 guang tou shan sui mi ji
Cardamine griffithii J. D. Hooker & Thomson var. grandifolia T. Y. Cheo & R. C. Fang.
Herbs perennial, 10-24(-30) cm tall, sparsely to densely pilose, often glabrescent above, rarely glabrous throughout. Rhizomes slender, with 1 or few filiform stolons. Stems erect, simple, terete, crisped pilose at base. Basal leaves not rosulate, often withered by anthesis; petiole 0.5-2 cm; terminal leaflet reniform or suborbicular, 0.5-1.5 × 0.7-1.7 cm, base cordate, margin shallowly crenate or repand; lateral leaflets 2, much smaller than terminal ones, sometimes absent and leaf simple. Middle and upper cauline leaves trifoliolate, sessile, to 10 cm; terminal leaflet reniform, suborbicular, or broadly ovate, 1.5-6(-7) × 1.2-4(-5) cm, with a petiolule 0.3-2.5(-5.2) cm, base cordate, rounded, or cuneate, glabrous, ciliate, or trichomes forming small tufts at teeth, margin crenate, repand, or obscurely to distinctly 2-6-toothed on each side, apex obtuse or acute; lateral leaflets 2, auriclelike, attached at or just above node, often giving appearance of amplexicaul leaf base, suborbicular, ovate, obovate, or reniform, 3-16(-25) × 2-13(-17) mm, much smaller than terminal leaflet, margin entire, 1- or 2-toothed, or repand. Racemes 3-10-flowered. Fruiting pedicels 0.5-1.7 cm, divaricate or ascending, slender. Sepals ovate, 2.5-3 × 1-1.5 mm, lateral pair subsaccate. Petals white, obovate, 6-8 × 2-3 mm, apex rounded. Median filament pairs 3-4.5 mm; lateral pair 1.5-3 mm; anther oblong, 0.7-1 mm. Ovules 8-16 per ovary. Fruit linear, slightly flattened, 1.5-2.5 cm × 1-1.2 mm; valves smooth, glabrous; style 1-2 mm. Seeds brown, oblong, 1.5-1.8 × 0.8-1 mm, wingless. Fl. and fr. Apr-Jul.
* Shady slopes, woodlands, moist places in ravines; 800-2900 m. Anhui, Fujian, Gansu, Hubei, Hunan, Shaanxi, Sichuan.
The above first record from Anhui is based on K. Yao 9028 (MO).