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FOC | Family List | FOC Vol. 8 | Brassicaceae

51. Crucihimalaya Al-Shehbaz et al., Novon. 9: 298. 1999.

须弥芥属 xu mi jie shu

Crucihimalaya himalaica

Credit: Photo: Al-Shehbaz

Herbs annual or biennial, rarely perennial with a caudex. Trichomes simple and stalked, 1- or 2-forked, sometimes stellate. Stems erect or ascending. Basal leaves rosulate or not, entire or dentate, rarely pinnately lobed. Cauline leaves sessile or subsessile, usually auriculate or sagittate, entire, dentate, or rarely pinnately lobed, rarely absent. Racemes several to many flowered, ebracteate or bracteate. Fruiting pedicels erect, ascending, or divaricate. Sepals oblong, erect, pubescent, base of lateral pair not saccate. Petals white, pink, or purple, longer than sepals; blade spatulate, rounded; claw obscurely distinct from blade. Stamens 6, erect, slightly tetradynamous; filaments not dilated at base; anthers ovate or oblong, obtuse at apex. Nectar glands confluent and subtending bases of filament. Ovules (30-)40-120(-150) per ovary. Fruit dehiscent siliques, linear, terete or somewhat 4-angled, rarely latiseptate, sessile or subsessile; valves with a distinct midvein, glabrous or rarely pubescent, smooth or torulose; replum rounded; septum complete; style to 1 mm; stigma capitate, entire. Seeds uniseriate, wingless, oblong, plump; seed coat minutely reticulate, mucilaginous when wetted; cotyledons incumbent.

Nine species: C and SW Asia, Himalayas, Mongolia, Russia; six species in China.

1 Fruit valves densely and coarsely stellate; pedicels pubescent all around; fruit often subappressed to rachis   2 C. lasiocarpa
+ Fruit valves glabrous or very rarely puberulent; pedicels glabrous adaxially, rarely (C. wallichii) pubescent all around; fruit not appressed to rachis.   (2)
2 (1) Cauline leaves distinctly auriculate or amplexicaul at base.   (3)
+ Cauline leaves petiolate or sessile, neither auriculate nor amplexicaul, sometimes absent.   (4)
3 (2) Racemes bracteate at least along proximal portion; annuals or biennials; cauline leaves usually auriculate; upper leaves coarsely stellate   4 C. himalaica
+ Racemes ebracteate; perennials; cauline leaves sagittate or amplexicaul; upper leaves softly stellate   1 C. mollissima
4 (2) Lowermost flowers of main raceme ebracteate; basal leaves lyrate or pinnatifid, often canescent, persistent in flower and fruit   6 C. wallichii
+ Lowermost flowers of main raceme bracteate; basal leaves entire or dentate, not canescent, withered by flowering or fruiting.   (5)
5 (4) Cauline leaves adaxially with simple and forked trichomes, ovate, elliptic, or oblong, rarely oblanceolate; plants 4-15(-20) cm tall; main raceme bracteate nearly throughout   5 C. axillaris
+ Cauline leaves adaxially with stellate stalked trichomes, linear-lanceolate; plants (18-)30-85(-120) cm tall; only lowermost flowers of main raceme bracteate   3 C. stricta

Lower Taxa


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