3. Adenia heterophylla (Blume) Koorders, Exkurs.-Fl. Java. 2: 637. 1912.
异叶蒴莲 yi ye shuo lian
Modecca heterophylla Blume, Bijdr. 940. 1826; Adenia chevalieri Gagnepain; A. maclurei Merrill; A. parviflora (Blanco) Cusset; A. populifolia K. Schumann & Lauterbach (1900), not Engler (1891); Modecca formosana Hayata; Passiflora parviflora Blanco.
Canopy lianas, to 30 m long. Roots fleshy, thickened. Stems terete, to 5 cm in diam.; internodes 2-20 cm; tendrils (3-forked or) simple, to 25 cm. Stipules flattened-triangular, ca. 1 mm; petiole 1-10 cm, glands at blade-petiole junction 2, on concave 2-4 mm in diam. auriculate extensions at apex of petiole in mature leaves, 1-3 mm in diam.; leaf blade ovate to lanceolate, membranous to leathery, glands often present, dotlike, base of mature leaf oblong, unlobed or (2 or)3-lobed with distal 1/3 of leaf often slightly constricted from slight lobation, margin entire to serrate, apex acute to acuminate. Plants (monoecious or) dioecious. Inflorescences often with tendril emerging from center, peduncle to 20 cm, to 40-flowered in males, 2-4-flowered in females; bracts and bracteoles narrowly triangular, ca. 1 mm, apex acute. Pedicel 0.5-13 mm. Flowers tubiform-urceolate, 15-25 × 2-5 mm, including stipe 1-15 mm. Hypanthium urceolate to cupuliform, wall fleshy-leathery, ca. 0.5 mm thick, 3-6 mm deep. Calyx tube 2-3 mm; lobes reflexed, triangular, 1-3 mm, rounded to acute, entire. Petals oblanceolate to triangular, 2-4 × 0.5-1 mm, red punctate, base hastate, apex entire to undulate, rounded to acute. Septa 0.25-3 mm high. Corona absent. Disk glands linear, to 3 mm high, truncate. Male flowers: filaments 1-4 mm, connate up to 2 mm, inserted at base of hypanthium; anthers ca. 3 × 1 mm, acute, up to 0.5 mm apiculate; vestigial ovary ca. 1 mm, gynophore to 1 mm. Female flowers: staminodes 1-3 mm, connate to 1 mm, inserted at base of hypanthium; ovary subglobose to oblong, ca. 4 × 2.5 mm; gynophore ca. 3 mm; style arms ca. 1 mm, free or connate for ca. 0.5 mm; stigmas globose, papillate. Capsules 1-3 per inflorescence, outside deep red at dehiscence, ellipsoid, fruit body to 13 cm, with gynophore to 3 cm, fruit wall leathery. Seeds to 60 per capsule, orbicular; funicles of seeds 5-15 mm. Fl. and fr. throughout year, primarily during wet periods.
Forests, scrub; near sea level to 300(-1000) m. Guangdong, Guangxi, Hainan, Taiwan [Cambodia, Indonesia (Java, Sumatra), Laos, New Guinea, Philippines, Thailand, Vietnam; Australia, Pacific islands].
This is a widespread and highly variable species. It has been treated as Adenia heterophylla [subsp. heterophylla] var. heterophylla according to de Wilde (Meded. Landbouwhogeschool Wageningen 71(18): 216. 1971).