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FOC | Family List | FOC Vol. 20-21 | Asteraceae | Klasea

8. Klasea centauroides (Linnaeus) Cassini ex Kitagawa, J. Jap. Bot. 21: 138. 1947.

麻花头 ma hua tou

Herbs (10-)25-150 cm tall. Stem erect, simple or branched, sparsely to densely crispate-pubescent near base, subglabrous in upper part, base with lacerate remains of old petioles. Leaves subglabrous to scabrid; pubescence often sparse, somewhat denser toward margin and along veins, composed of small appressed (ca. 0.2 × 0.03 mm) and larger patent (ca. 1 × 0.1 mm) multicellular hairs. Basal and lower stem leaves petiolate. Upper leaves sessile. Capitula 1 to many. Involucre 0.6-3 cm in diam. Phyllaries imbricate, in 4-10 rows; outer and middle phyllaries 2-11 × 1.5-5(-6) mm, without an apical spine or with a 0.2-2 mm spinule. Corolla purple to pink, rarely almost white, 2-3 cm. Achene brown to pale brown, cuneate-ellipsoid to obliquely cuneate, 4.5-6 mm, faintly ribbed. Pappus 7-11 mm. Fl. and fr. May-Oct.

Mountain slopes, steppes, meadows, sandy hills, open forests, forests, grasslands, wastelands, roadsides, riversides, watersides, farmlands; 200-3500 m. Anhui, Gansu, Hebei, Heilongjiang, Henan, Jilin, Liaoning, Nei Mongol, Ningxia, Qinghai, Shaanxi, Shandong, Shanxi, N Sichuan [Korea, Mongolia, Russia].

Klasea centauroides is a very polymorphic species with respect to number of capitula, leaf shape, shape of involucre, and coloration of phyllaries. Several distinctive forms have been described as species, but all of them are connected by intermediate forms. Delimitation of subspecies is problematic especially in regions where their ranges overlap. Herbarium specimens are often misidentified; therefore, distribution data from literature have to be taken with care and may be incomplete for some subspecies treated here.


1 Stem much branched; capitula (3-)10-60; involucre 0.6-1.2 cm in diam.   8b subsp. polycephala
+ Stem simple or sparingly branched; capitula 1-6(-12); involucre 1.1-3 cm in diam   (2)
       
2 (1) All leaves undivided.   8e subsp. cupuliformis
+ At least middle stem leaves pinnatisect or pinnatipartite   (3)
       
3 (2) Involucre obconic to bowl-shaped; inner phyllaries straight, without clear distinction between basal part and scarious appendage; middle phyllaries yellowish green with pale margin.   8f subsp. chanetii
+ Involucre semiglobose, ovoid, or campanulate; inner phyllaries mostly geniculate between basal part and appendage; middle phyllaries with dark tip and/or purple scarious margins   (4)
       
4 (3) Involucre base shortly attenuate; middle phyllaries 1.8-3 mm wide; lateral segments of middle stem leaves usually antrorse at an acute angle.   8c subsp. komarovii
+ Involucre base rounded or slightly umbilicate; middle phyllaries 2.5-6 mm wide; lateral segments of middle stem leaves usually at right angle or nearly so   (5)
       
5 (4) Stem ± foliate throughout; peduncles 1-12(-18) cm; involucre not or slightly constricted, 1.1-2.2 cm in diam.   8a subsp. centauroides
+ Stem ± leafless in upper part; peduncles (10-)15-40 cm; involucre conspicuously constricted, (1.5-)2-3 cm in diam.   8d subsp. strangulata

Lower Taxa


 

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