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Pakistan | Family List | Ranunculaceae

14. Clematis L., Sp. Pl. 543. 1753. Gen. Pl. ed. 5:616. 1754, Boiss., Fl. Or. 1:2. 1867, Hook.f. & Thoms. in Hook.f, Fl. Brit. Ind. 1:2. 1872, O. Kuntze, Verh. Dot. Ver. Brandenburg 26: 83-202. 1881 & Verh. Zool. Bot. Ges. Wien 37: 47-50. 1887, Qureshi & Chaudhri in Pak. Syst. 4(1-2):121.1988.

HARALD RIEDL and YASIN J. NASIR


Naturhistorisches Museum, Botanische Abteilung, Wien, Austria.

and

National Herbarium, Pakistan Agricultural Research Council, Islamabad, Pakistan.

Clematis montana
Illustration

Climbing to erect shrubs, perennials rarely annuals. Leaves opposite, ternately or pinnately divided. Flowers solitary (-2-4) in fascicles or in paniculate cymes. Sepals 4-8, petaloid. Petals absent but filaments or staminodia sometimes petaloid. Stamens numerous. Carpels forming a small subglobose head. Ovules solitary, pendulous. Style often much elongating after flowering, plumose-hairy.

A genus of c. 250 cosmopolitan, but chiefly temperate species. Species with petaloid filaments or staminodes are sometimes separated in subgenus Atragene L.

Several species as Clematis hybrida Hort. and Clematis flammula L. are cultivated in gardens for their ornamental value. No poisonous plants are known in Clematis. Represented in Pakistan by 10-11 species.


1 Leaves pinnate, with very small lateral leaflets and much longer, simple, terminal leaflets   7 Clematis aspleniifola
+ Leaves ternate to pinnate with ± equal leaflets   (2)
       
2 (1) Leaflets linear-lanceolate to narrowly ovate, never cordate at base, sometimes distinctly lobed with entire lobes   (3)
+ Leaflets with cordate base or ± broadly ovate, grossly dentate, rarely entire or lobed with a few grossly dentate lobes   (6)
       
3 (2) Flowers single on long pedicels. Corolla-l ike sepals yellow, at least 3 cm long   (4)
+ Flowers in panicles or few-flowered fascicles, not distinctly corolla-l ike, sepals white or pale yellow, less than 3 cm   (5)
       
4 (3) Filaments dilated, petaloid, but always bearing fertile anthers   1 Clematis robertsiana
+ Petaloid staminodes present   10 Clematis alpina var. sibirica
       
5 (3) Filaments longer than anthers   8 Clematis graveolens
+ Filaments shorter than anthers   9 Clematis orientalis
       
6 (2) Leaves ternate. Flowers single or in fascicles on long pedicels   (7)
+ Leaves pinnate or bi-pinnate   (8)
       
7 (6) Sepals white to yellowish-white, obtuse to subacute, oblong to ovovate. Anthers glabrous   2 Clematis montana
+ Sepals violet outside, narrower, acute to sub-acute. Anthers slightly bearded   11 Clematis barbellata
       
8 (6) Leaflets leathery, ± riged, often shiny on the upper surface. Flowers densely paniculate   3 Clematis gouriana
+ Leaflets neither leathery nor rigid   (9)
       
9 (8) Sepals 6-8 (-10) mm long, horizontally spreading   4 Clematis grata
+ Sepals 10-20 mm long-erect, curved backwards at the tips   (10)
       
10 (9) Sepals with ± parallel ribs, 15-20 mm long   5 Clematis buchananiana
+ Sepals without ribs, 10-12 mm long   6 Clematis connata

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