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Pakistan | Family List | Boraginaceae

30. Heterocaryum A. DC. in DC., Prodr. 10:144. 1846. Brand in Engler, Pflanzenr. iv. 252:95. 1931; Riedl in Rech. f., Fl. Iran. 48.84.1967; Kazmi in J. Arn. Arb. 51(3): 515.1970.


Annual medium-sized herbs. Leaves narrow. Pedicels long, usually thickened. Calyx divided to the base, somewhat enlarging in fruit. Corolla small, 5-lobed. Throat appendages present. Style short, stigma capitate. Nutlets attached to a gynobase throughout their length and not detachable. Gynobase columnar. Nutlets 4, usually unequal, compressed, margin winged or not-if winged the dorsal middle area smooth; appendages glochidiate. Closely related to Lappula Gilib. and sometimes united with it.

6 species distributed from S. Russia & Turkey to C. Asia and Himalaya. Represented in Pakistan by 3 species.

1 Plants with subglabrous parts. Fruiting pedicel slender, 0.8-1 mm thick, often strongly reflexed. Wing of nutlets with appendages in two series   1 Heterocaryum laevigatum
+ Plants with subpatent to appressed hairs, sometimes densely so. Fruiting pedicel if present, stout, 1.5-2 mm thick, erect to slightly recurved   (2)
2 (1) Fruiting pedicel absent or very short, 1-1.5 mm long   2 Heterocaryum subsessile
+ Fruiting pedicel up to 10 mm long   3 Heterocaryum szovitsianum

Lower Taxa

Related Synonym(s):


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