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Pakistan | Family List | Guttiferae

Hypericum L., Sp.P1. 783. 1753. Gen. Pl.ed.341 .1 754 ; W.T.Thiselton Dyer in Hook.f., F1.Brit.Ind.l:253.1874; H.Collett, Fl. Simlensis: 54.1902; R. Keller in Engler and Prantl, Nat. Pflanzenfam., ed. 2.25:175.1925 pro parte excl. Sect IV Elodea; N.K.B. Robson in P.H. Davis, Fl. Turkey, 2:355.1966; in Rech. f., Fl. Iranica, 49, Guttiferae: 1. 1968.

Hypericum napaulense

Credit: V.G.

Trees, shrubs or perennial to annual herbs, glabrous (in W. Pakistan species), with translucent, red or black glands. Leaves opposite, entire or rarely gland-fringed. Flowers solitary or in terminal dichasial to monochasial cymes or panicles. Sepals 5, quincuncial or rarely 4, decussate, glandular like the leaves. Petals 5(4), yellow, often tinged or veined red, deciduous or persistent. Stamens fascicles 5(4), antepetalous, free or variously united (often 2+2+1, forming `3'fascicles), deciduous or persistent; filaments slender, free or united; anthers dorsifixed, with an amber or black gland on the connective. Ovary 1-5-locular, with (2)3-5 placentae; styles (2)3-5, free or ± united, slender; ovules 2 to numerous on each placenta. Fruit a septicidal capsule or rarely tricoccoid or indehiscent. Seeds sometimes winged or carinate, with variously patterned testa.

A genus of c. 400 species, almost cosmopolitan, but absent from arctic regions and rare in Australasia (2 species) and lowland tropical regions.

1 Stamen fascicles and styles 5. petals and stamens deciduous. Shrubs. Black glands absent.   (2)
+ Stamen fascicles and styles 3 petals and stamens persistent. Herbs. Black glands present   (6)
2 (1) Styles free, sometimes mutually adpressed at the base when young (native species)   (3)
+ Styles united, almost to the apex (cultivated)   Hypericum monogynum
3 (2) Styles 1.5-3 x ovary, erect or gradually divergent. Petals narrowly obovate to oblanceolate, 1.2-1.5 x stamens   (4)
+ Styles 0.3-1.0 x ovary, outcurving. Petals broadly obovate to subcircular, 2-4 x stamens   (5)
4 (3) Sepals ovate to elliptic-oblong, erect in bud. Flowers 3.5-7.5 cm in diam. Leaves sessile, oblong or ovate-oblong to elliptic   1 Hypericum oblongifolium
+ Sepals linear or narrowly oblong-lanceolate, spreading in bud. Flowers 1.5-3.5 cm in diam. Leaves shortly petiolate, ovate to lanceolate or elliptic-lanceolate   Hypericum dyeri
5 (3) Sepals acute to subacute at apex, ovate to lanceolate or narrowly elliptic, often foliaceous. Branches 4-lined at first, soon terete, not markedly frondose   Hypericum choisianum
+ Sepals rounded at apex, oblong or elliptic to obovate-spathulate, not foliaceous. Branches 4-angled or 4-lined at first, later 2-lined or (rarely) terete, usually + markedly frondose   Hypericum uralum
6 (1) Stem + scabrid. Inflorescence corymbose, dense. Black glands present on petal and or sepal margins only   Hypericum scabrum
+ Stem smooth. Inflorescence paniculate or few-flowered. lax. Black glands present on anthers and leaves and fre-quently also on stem, sepals and petasl   (7)
7 (6) Stem 2-lined with black glands on raised lines. Sepals entire. Capsule valves with longitudinal dorsal vittae and lateral vesicles   Hypericum perforatum
+ Stem usually terete, eglandular. Sepals glandular-ciliate to glandular-fimbriate (in W. Pakistan). Capsule valves with longitudinal vittae only   (8)
8 (7) Petals with marginal black glands. Styles 1.5-2.5 x longer than ovary. Stem always terete, usually unbranched. Leaves acute to obtuse. Sepals ribbed   Hypericum elodeoides
+ Petals without marginal black glands. Styles 0.9-1.3 x as long as ovary. Stem sometimes 2-6-lined, frequently bran- ched. Leaves subobtuse to rounded. Sepals not or scarcely ribbed   9 Hypericum napaulense

Lower Taxa

Related Synonym(s):


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