1 |
Leaves reniform or orbicular-reniform, cordate-reniform, or hastate, palmately or palmate-pinnately veined |
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(2) |
+ |
Leaves variable in shape, pinnately or parallel veined |
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(21) |
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2 (1) |
Capitula without ray florets; phyllaries apically rounded or obtuse |
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(3) |
+ |
Capitula with ray florets; phyllaries apically acute or acuminate |
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(7) |
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3 (2) |
Phyllaries petaloid, membranous, purplish red or rarely yellowish white, broadly elliptic or obovate-oblong, outside purplish red pilose, rarely white pilose, apex rounded |
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(4) |
+ |
Phyllaries leaflike, herbaceous, dark brown, oblong or oblanceolate, outside glabrous or pilose, apex obtuse |
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(5) |
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4 (3) |
Leaf margin regularly crenate or lobed. |
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1 C. campanulatum |
+ |
Leaves palmatisect. |
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2 C. pinnatisectum |
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5 (3) |
Phyllaries oblong, margin broadly brown membranous, apex entire; pappus shorter than tubular corolla; leaves glabrous, abaxially pale green, adaxially dark green. |
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3 C. calcicola |
+ |
Phyllaries usually oblanceolate, margin not membranous, apex denticulate or erose; pappus as long as tubular corolla; leaves concolorous on both surfaces |
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(6) |
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6 (5) |
Sheaths of stem leaves tubular, outside glabrous; phyllaries black, base shortly pilose, outside glabrous; pappus white. |
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4 C. atrocapitatum |
+ |
Sheaths of stem leaves cucullate, outside white arachnoid-puberulent; phyllaries outside glabrous; pappus yellowish brown. |
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5 C. cucullatum |
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7 (2) |
Ray lamina yellow, oblong to lanceolate, apex acuminate, rarely obtuse |
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(8) |
+ |
Ray lamina purplish red or yellow, broadly oblanceolate or cuneate, apex truncate or rounded, 3-lobed, rarely 3-denticulate |
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(12) |
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8 (7) |
Ray lamina oblong, apex acute or obtuse |
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(9) |
+ |
Ray lamina ovate- or linear-lanceolate, apex acuminate or caudate |
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(10) |
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9 (8) |
Leaves glabrous, with conspicuous white reticulate veins on both surfaces or at least abaxially; involucre outside initially shortly pilose, glabrescent; ray lamina apically obtuse. |
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6 C. smithianum |
+ |
Leaves abaxially brown pilose, without reticulate veins; involucre outside brown pilose; ray lamina apically acute. |
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7 C. decaisnei |
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10 (8) |
Ray lamina linear-lanceolate, membranous, transparent; pappus as long as tubular corolla; leaves angular dentate. |
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8 C. stenoglossum |
+ |
Ray lamina ovate-lanceolate, not transparent; pappus shorter than to as long as tubular corolla; leaves crenate or coarsely dentate |
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(11) |
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11 (10) |
Pappus brown, shorter than tubular corolla; leaves green on both surfaces, margin crenate. |
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9 C. citriflorum |
+ |
Pappus yellowish brown, as long as tubular corolla; leaves abaxially pale green, adaxially green, margin coarsely dentate. |
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10 C. pulchrum |
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12 (7) |
Ray lamina purple; pappus white |
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(13) |
+ |
Ray lamina yellow; pappus white or brown |
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(15) |
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13 (12) |
Leaves palmatipartite; tubular florets purplish red. |
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11 C. palmatum |
+ |
Leaf margin regularly crenate; tubular florets yellow or purplish red |
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(14) |
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14 (13) |
Leaves abaxially densely white arachnoid-pilose; tubular florets yellow; styles proximally yellowish white, distally blackish gray, as long as tubular corolla. |
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12 C. farreri |
+ |
Leaves abaxially sparsely pilose; tubular florets purplish red, styles purplish red, 2-2.5 cm, longer than tubular corolla. |
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13 C. rhodocephalum |
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15 (12) |
Leaves cordate-reniform or hastate |
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(16) |
+ |
Leaves reniform or orbicular-reniform |
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(18) |
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16 (15) |
Stem simple; leaves hastate, leaf margin usually entire or sparsely irregularly dentate; pappus as long as tubular corolla. |
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18 C. forrestii |
+ |
Stems several; leaves cordate-reniform to hastate, leaf margin dentate; pappus shorter than tubular corolla |
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(17) |
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17 (16) |
Leaves cordate-reniform; involucre ca. 20 mm; pappus brown, ca. 4 mm, much shorter than tubular corolla. |
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19 C. medogense |
+ |
Leaves cordate-reniform to hastate; involucre 30-32 mm; pappus yellowish, 7-8 mm, slightly shorter than tubular corolla. |
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20 C. latilobum |
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18 (15) |
Plants slender; leaves thin, margin angular dentate; ray lamina apically rounded, denticulate; pappus white. |
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14 C. reniforme |
+ |
Plants robust; leaves thick, margin regularly dentate; ray lamina apically truncate, 3- or 4-lobed |
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(19) |
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19 (18) |
Stem leaves with bulbil inside sheaths; pappus white. |
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15 C. bulbilliferum |
+ |
Stem leaves without bulbil inside sheaths; pappus brown |
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(20) |
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20 (19) |
Basal leaves glabrous; petiole glabrous or distally brown pilose; petiole of stem leaves enlarged into leaf blade, margin sharply dentate; pappus shorter than tubular corolla. |
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16 C. phyllodineum |
+ |
Basal leaves abaxially shortly black pilose along veins; petiole glabrous; petiole of stem leaves sheathlike, margin entire; pappus as long as tubular corolla. |
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17 C. thomsonii |
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21 (1) |
Leaves bluish green or grayish green, linear to broadly elliptic, parallel or straight veined |
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(22) |
+ |
Leaves green or differently colored on both surfaces, lanceolate to oblong, pinnately veined |
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(27) |
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22 (21) |
Involucre outside densely brown or purplish brown pilose; leaves lanceolate to elliptic |
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(23) |
+ |
Involucre outside glabrous; leaves linear to oblanceolate |
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(25) |
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23 (22) |
Capitulum solitary, without ray florets. |
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21 C. angustifolium |
+ |
Capitula 1-13, in raceme or solitary, with ray florets; lamina membranous, transparent, linear-lanceolate, 25-70 × 1.5-5 mm, apex acuminate |
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(24) |
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24 (23) |
Leaflike bracts linear to lanceolate, herbaceous, green. |
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22 C. brunneopilosum |
+ |
Leaflike bracts ovate, membranous, yellowish white. |
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23 C. stenactinium |
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25 (22) |
Ray lamina linear-lanceolate, to 4 cm, apex acuminate or absent; leaves linear to linear-lanceolate, usually 2-5 mm wide, rarely to 30 mm wide. |
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24 C. lineare |
+ |
Ray lamina oblong, elliptic, or oblanceolate, 12-16 × 3-6 mm, apex obtuse or truncate; leaves oblong or oblanceolate to linear, 2-11 mm wide |
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(26) |
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26 (25) |
Leaves linear, 2-3.5 mm wide, margin revolute, entire, apex rounded. |
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25 C. bhutanicum |
+ |
Leaves oblong or oblanceolate, 6-11 mm wide, margin remotely denticulate, apex acute. |
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26 C. conaense |
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27 (21) |
Ray lamina broadly oblanceolate or cuneate, apex truncate, lobed |
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(28) |
+ |
Ray lamina elliptic or lanceolate to linear-lanceolate, apex acute or acuminate |
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(31) |
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28 (27) |
Leaves pinnatipartite to pinnatisect |
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(29) |
+ |
Leaf margin entire or dentate, rarely lobed |
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(30) |
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29 (28) |
Leaves ovate, lobes narrowly lanceolate, 10-30 × 1-4 mm; involucre glabrous. |
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27 C. dissectum |
+ |
Leaves oblong, lobes oblong, 4-8 × ca. 1 mm; involucre blackish pilose. |
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28 C. pinnatifidum |
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30 (28) |
Leaves spatulate, margin triangularly dentate or denticulate, base truncate; pappus white; stem densely white pilose. |
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29 C. sino-oblongatum |
+ |
Leaves hastate or ovate-cordate, margin entire to lobed or crenate, base cordate; pappus pale brown; stem glabrous. |
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30 C. trilobum |
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31 (27) |
Plants greenish gray or bluish green, glabrous, usually mealy; stem leaves numerous, erect, adnate, tubular-amplexicaul or only base amplexicaul |
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(32) |
+ |
Plants green, hairy, at least stem distally and involucre at base, rarely glabrous |
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(41) |
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32 (31) |
Leaflike bracts as leaf blade, usually surrounding involucre; ray lamina narrowly lanceolate, apex acuminate |
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(33) |
+ |
Leaflike bracts linear, not surrounding involucre |
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(34) |
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33 (32) |
Leaves ovate-elliptic to broadly elliptic, base cuneate. |
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31 C. helianthus |
+ |
Leaves narrowly elliptic or spatulate, base gradually narrowed. |
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32 C. suave |
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34 (32) |
Capitula 5-10, in raceme; ray lamina 5-7 mm, slightly longer than involucre; pappus pale yellow, 2-3 mm, ca. 1/2 as long as tubular corolla |
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(35) |
+ |
Capitulum solitary; ray lamina longer than involucre; pappus white or pale yellow |
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(36) |
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35 (34) |
Basal leaves broadly elliptic, base gradually narrowed into a narrowly winged petiole; stem leaves apically obtuse. |
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33 C. botryocephalum |
+ |
Basal leaves broadly spatulate-oblong, base gradually narrowed into a broadly winged petiole; stem leaves apically acuminate. |
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34 C. spathulifolium |
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36 (34) |
Leaves petiolate; petiole 3-11 cm, slender; leaf blade oblong or elliptic, base rounded |
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(37) |
+ |
Leaves sessile or shortly petiolate; leaf blade ovate or lingulate-spatulate; ray lamina apically acuminate; pappus white, as long as tubular corolla |
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(39) |
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37 (36) |
Pappus white, as long as tubular corolla; ray lamina apically acuminate. |
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35 C. glaucum |
+ |
Pappus much shorter than tubular corolla or absent; ray lamina apically obtuse, acute, or acuminate |
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(38) |
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38 (37) |
Pappus absent; ray lamina apically acuminate. |
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36 C. bupleurifolium |
+ |
Pappus pale yellow, 1-2 mm, 1/5-1/2 as long as tubular corolla; ray lamina apically acute or obtuse. |
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37 C. brachychaetum |
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39 (36) |
Leaves lingulate-spatulate, base gradually narrowed into a broad petiole. |
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38 C. lingulatum |
+ |
Leaves ovate or cordate, base cordate, narrowly petiolate |
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(40) |
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40 (39) |
Leaves ovate; petiole glabrous. |
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39 C. yadongense |
+ |
Leaves cordate; petiole brown pilose. |
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40 C. cordatum |
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41 (31) |
Phyllaries in 1 row, equal in size, base united, cupular |
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(42) |
+ |
Phyllaries in 2 rows, separate, outer narrow, inner broad, outside glabrous or hairy |
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(47) |
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42 (41) |
Leaves 12-15 × 6-9 cm; petiole 15-30 cm; capitula 3, in corymb-raceme, long pedunculate; involucre outside black pilose; ray lamina linear-lanceolate. |
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41 C. petiolatum |
+ |
Leaves 0.7-6 × 1-4 cm, shortly petiolate; capitulum solitary; involucre outside black or white pilose; ray lamina oblong |
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(43) |
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43 (42) |
Ray lamina as long as involucre or tubular florets; phyllaries apically rounded; leaves abaxially white pilose |
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(44) |
+ |
Ray lamina longer than involucre; phyllaries apically acute or acuminate; leaves glabrous or abaxially pilose |
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(45) |
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44 (43) |
Outer florets radiate; lamina yellow, as long as involucre. |
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42 C. nanum |
+ |
Outer florets radiate or narrowly tubular, as long as tubular florets in center; lamina white, smaller or absent; slender tubular florets with short corolla. |
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43 C. microglossum |
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45 (43) |
Plants with rosette leaves; underground stems absent; leaves glabrous, margin revolute. |
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44 C. microphyllum |
+ |
Plants without rosette leaves; leaves all stem, abaxially densely white pilose, adaxially glabrous; underground stem rhizomelike, with scale leaves |
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(46) |
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46 (45) |
Leaf margin entire to shallowly dentate. |
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45 C. humile |
+ |
Leaf margin pinnately laciniate; segments linear-oblong. |
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46 C. laciniatum |
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47 (41) |
Ray florets absent; involucre outside densely dark brown pilose; leaves abaxially greenish white, adaxially dark green, margin entire. |
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47 C. discoideum |
+ |
Ray florets present, if ray florets absent or with short lamina then involucre outside blackish gray pilose; leaf margin coarsely dentate |
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(48) |
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48 (47) |
Capitula large; ray lamina tapelike, narrowly lanceolate to linear-lanceolate, usually 2-4 × as long as involucre, apex acuminate |
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(49) |
+ |
Capitula small; ray lamina oblong, elliptic to linear, often 1-2 × as long as involucre |
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(56) |
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49 (48) |
Leaves densely hairy, oblong, obovate to orbicular; involucre outside white and black pilose |
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(50) |
+ |
Leaves glabrous, variable in shape; involucre outside blackish purple-brown or white pilose, rarely glabrous |
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(51) |
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50 (49) |
Leaves densely white pilose, base narrowed into a broadly winged petiole; capitula 1-4, solitary or in corymb-raceme. |
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48 C. obovatum |
+ |
Leaves densely shortly white pilose, base rounded, narrowly petiolate; capitulum solitary. |
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49 C. puberulum |
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51 (49) |
Leaf base cordate or truncate |
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(52) |
+ |
Leaf base cuneate |
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(54) |
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52 (51) |
Leaves ovate, base cordate, margin entire; ray lamina linear-lanceolate, 2-5 mm wide; pappus white. |
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50 C. prattii |
+ |
Leaves triangular or sagittate, margin dentate; ray lamina broadly lanceolate, 6-9 mm wide; pappus pale brown |
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(53) |
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53 (52) |
Leaves concolorous on both surfaces, triangular, 6-13 cm, base truncate. |
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51 C. delavayi |
+ |
Leaves abaxially pale green, adaxially dark green, sagittate, 1.5-3 cm, base cordate. |
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52 C. sagittifolium |
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54 (51) |
Involucre outside purplish red or white pilose; stem glabrous; leaves abaxially pale green, adaxially green, broadly lanceolate, margin coarsely dentate. |
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53 C. daochengense |
+ |
Involucre outside brown or dark brown pilose; stem shortly black pilose; leaves glabrous or yellow pilose, obovate or broadly elliptic, margin subentire or denticulate |
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(55) |
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55 (54) |
Leaves glabrous, margin subentire. |
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54 C. nobile |
+ |
Leaves yellow pilose, margin denticulate or subentire. |
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55 C. pilosum |
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56 (48) |
Capitula 2-13, in raceme; stem leaves normal; plants robust, tall |
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(57) |
+ |
Capitulum solitary, rarely 3; stem leaves bracteal, ovate-oblong to linear; plants dwarf |
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(59) |
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57 (56) |
Involucre outside black pilose; ray lamina broadly elliptic; stem leaves ovate. |
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56 C. arnicoides |
+ |
Involucre outside white pilose; ray lamina oblong; stem leaves oblong or ovate-oblong |
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(58) |
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58 (57) |
Leaves herbaceous, white arachnoid-puberulent, scarcely veined; stem leaves with sheath amplexicaul; peduncles equal, to 1 cm; raceme not spreading. |
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57 C. chungdienense |
+ |
Leaves leathery, glabrous, prominently veined; stem leaves not amplexicaul; peduncles unequal, 2-6 cm; raceme spreading. |
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58 C. coriaceum |
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59 (56) |
Pappus brown; leaf base cordate |
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(60) |
+ |
Pappus white; leaf base usually cuneate, truncate, or rounded, if base cordate then involucre outside shortly black pilose |
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(62) |
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60 (59) |
Basal leaves ovate-cordate or ovate-lanceolate to lanceolate; involucre broadly campanulate; ray lamina linear. |
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59 C. potaninii |
+ |
Basal leaves ovate-cordate to orbicular-cordate or squarelike; involucre hemispheric; ray lamina oblong |
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(61) |
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61 (60) |
Leaves small, leathery, nitid, conspicuously reticulate veined. |
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60 C. cyclaminanthum |
+ |
Leaves large, herbaceous, scarcely reticulate veined, abaxially glabrous or brown pilose along veins. |
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61 C. principis |
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62 (59) |
Involucre outside shortly black pilose or glabrous; leaf base cordate or rounded; ray lamina slightly longer than involucre |
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(63) |
+ |
Involucre outside white or blackish gray pilose, rarely glabrous; leaf base usually cuneate, rarely truncate |
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(65) |
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63 (62) |
Leaves thin, often purplish red, scarcely reticulate veined. |
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62 C. purpureifolium |
+ |
Leaves thick, green, conspicuously prominently reticulate veined |
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(64) |
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64 (63) |
Basal leaves reticulate veined on both surfaces; capitula sometimes 3; ray lamina broadly elliptic. |
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63 C. nervosum |
+ |
Basal leaves abaxially reticulate veined; capitulum solitary; ray lamina oblong. |
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64 C. nepalense |
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65 (62) |
Basal leaves narrowly petiolate; petiole densely pilose, not winged; stem leaves numerous, linear, erect, margin entire; involucre outside glabrous; ray lamina slightly longer than involucre, apex truncate. |
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65 C. variifolium |
+ |
Basal leaves broadly petiolate; petiole winged, glabrous or hairy, sometimes enlarged into a leaflike blade; stem leaves ovate-oblong to lanceolate, margin dentate; ray lamina 2-3 × as long as involucre, apex acute |
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(66) |
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66 (65) |
Involucre outside glabrous, phyllaries narrowly lanceolate, apex acuminate. |
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66 C. pseudo-oblongatum |
+ |
Involucre outside densely hairy, phyllaries lanceolate or ovate-lanceolate, apex acute or obtuse |
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(67) |
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67 (66) |
Involucre outside white arachnoid-puberulent. |
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67 C. oblongatum |
+ |
Involucre outside blackish gray pilose |
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(68) |
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68 (67) |
Leaves and petiole glabrous. |
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68 C. ellisii |
+ |
Leaves and petiole densely glandular pilose. |
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69 C. glandulipilosum |
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