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FOC | Family List | FOC Vol. 22 | Poaceae

50. Stipa Linnaeus, Sp. Pl. 1: 78. 1753.

针茅属 zhen mao shu

Perennials, forming dense tussocks, old basal sheaths persistent. Leaf blades filiform to setaceous, convolute, abaxial surface smooth or scabrid, adaxial surface prominently ribbed. Inflorescence usually a contracted panicle, enclosed in uppermost leaf sheath or shortly exserted, spikelets few. Spikelets with one floret, bisexual; glumes subequal, hyaline or membranous, much longer than floret, 3–5-veined, long acuminate; callus pungent, shortly bearded; lemma narrowly lanceolate, terete, usually leathery, (3–)5-veined, hairy, margins overlapping, apex entire; awn articulated at lemma apex and deciduous at maturity, scabrid to plumose, 1–2-geniculate, column tightly twisted, bristle straight, flexuous or curling; palea subequaling lemma, hyaline, enclosed within lemma. Lodicules 2 or 3, lanceolate. Stamens 3, anthers glabrous or shortly hairy at apex. Stigmas 2.

About 100 species: temperate and warm-temperate regions of Asia and Europe, in dry, open habitats; 23 species (three endemic) in China.


1 Awn scabrid or minutely hispidulous (hairs to 0.3 mm) throughout   (2)
+ Awn plumose or at least shortly hairy along part or all of its length   (8)
       
2 (1) Cleistogamous spikelets present in basal leaf sheaths; lemma 0.5–0.6 cm.   1 S. bungeana
+ Cleistogamous spikelets not present in basal leaf sheaths; lemma 0.8–1.7 cm   (3)
       
3 (2) Panicle branches not enclosed by uppermost leaf sheath, usually clearly exserted; awn minutely hispidulous, bristle straight or slightly flexuous   (4)
+ Panicle branches enclosed by expanded uppermost leaf sheath; awn scabrid, bristle curling   (5)
       
4 (3) Glumes 1.2–1.5 cm, apex finely acuminate; awns separate, bristle stiff.   2 S. przewalskyi
+ Glumes 1.5–3 cm, apex filamentous; awns twisted together at top of panicle, bristle flexuous.   3 S. capillacea
       
5 (3) Glumes 1.8–2.5 cm; awn column 1.5–2.5 cm to first bend.   4 S. sareptana
+ Glumes 2.5–4.5 cm; awn column 3–10 cm to first bend   (6)
       
6 (5) Ligule of culm leaves 1.5–2 mm.   5 S. baicalensis
+ Ligule of culm leaves 3–10 mm   (7)
       
7 (6) Lemma 1–1.2 cm; awn 13–18 cm.   6 S. capillata
+ Lemma 1.5–1.7 cm; awn 20–28 cm.   7 S. grandis
       
8 (1) Awn plumose, or at least shortly hairy, throughout its length   (9)
+ Awn plumose only on column or only on bristle   (16)
       
9 (8) Awn 1-geniculate (sometimes weakly 2-geniculate in S. consanguinea)   (10)
+ Awn 2-geniculate   (12)
       
10 (9) Awn shortly pubescent, hairs on bristle 0.4–0.7 mm.   9 S. consanguinea
+ Awn plumose, hairs on bristle 2–5 mm   (11)
       
11 (10) Panicle shortly exserted; lemma evenly hairy.   10 S. mongolorum
+ Panicle branches enclosed by uppermost leaf sheath; lemma hairs in lines.   11 S. caucasica
       
12 (9) Lemma hairs in lines; glumes gray-green or pale brownish, narrowly lanceolate, long attenuate-filiform   (13)
+ Lemma evenly hairy; glumes green or dark purple, lanceolate, acuminate into awn-point   (15)
       
13 (12) Glumes 1.8–3 cm; lemma smooth or scaberulous below apex; awn hairs 1–4 mm   (14)
+ Glumes 1.2–2 cm; lemma spinulose below apex; awn hairs 0.8–1.5 mm.   14 S. breviflora
       
14 (13) Culms 15–35 cm; awn 4–6 cm, hairs on bristle 3–4 mm.   12 S. orientalis
+ Culms 40–80 cm; awn 9–20 cm, hairs on bristle 1–2.5 mm.   13 S. arabica
       
15 (12) Panicle branches 3–6 cm, flexuous; glumes dark purple; awn 6.5–9 cm, hairs shorter below, 2–3 mm on bristle.   15 S. purpurea
+ Panicle branches 2–4 cm, straight; glumes green or purplish; awn 3–6 cm, hairs longer below, 0.3–0.7 mm on bristle.   16 S. roborowskyi
       
16 (8) Awn 1.5–3 cm; column plumose, bristle glabrous or minutely hairy (hairs 0.1–0.2 mm)   (17)
+ Awn 6–28 cm; column glabrous, bristle plumose, hairs 2–6 mm   (20)
       
17 (16) Panicle 1–2 cm wide; branches 1–3 cm, suberect   (18)
+ Panicle (2–)3–7 cm wide; branches 3–6 cm, spreading or obliquely ascending   (19)
       
18 (17) Awn obscurely 2-geniculate, hairs on column up to 1 mm; spikelets 1.1–1.4 cm.   17 S. regeliana
+ Awn distinctly 2-geniculate, hairs on column 2–3 mm; spikelets 0.6–1.1 cm.   18 S. subsessiliflora
       
19 (17) Leaf blades scabrid or hairy; panicle branches spreading, with cushion in axils; ligule 3–7 mm; column hairs 3–4 mm.   19 S. penicillata
+ Leaf blades smooth, glabrous; panicle branches ascending, without cushion in axils; ligule 1–1.5 mm; column hairs 1–2 mm.   20 S. aliena
       
20 (16) Awn 1-geniculate, 6–13 cm.   8 S. tianschanica
+ Awn 2-geniculate, 15–28 cm   (21)
       
21 (20) Glumes 2–3 cm; lemma evenly hairy; awn hairs 2–3 mm.   21 S. lessingiana
+ Glumes 3–5.5 cm; lemma hairs in lines; awn hairs 3–6 mm   (22)
       
22 (21) Column of awn 5–6 cm to first bend; ligule up to 4 mm.   22 S. kirghisorum
+ Column of awn 3.5–4.5 cm to first bend; ligule 4–12 mm.   23 S. macroglossa

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