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FOC | Family List | FOC Vol. 8 | Saxifragaceae | Saxifraga

4. Saxifraga sect. Ciliatae Haworth, Misc. Nat. 160. 1803.

山羊臭组 shan yang xiu zu

Hirculus Haworth.

Herbs mostly perennial, some possibly annual, biennial, or monocarpic perennial (suspected rather than demonstrated), forming erect individuals, cespitose clumps, cushions, or sprawling mats, lacking basal bulbils. Leaves alternate, very rarely opposite, petiolate or not; leaf blade herbaceous or carnose, lacking chalk glands and crystals, margin usually entire, occasionally toothed or lobed. Flowering stem usually leafy; inflorescence cymose or flower solitary. Flowers sometimes unisexual, actinomorphic, 5-merous. Petals usually yellow or orange, occasionally white, pink, or red, sometimes spotted orange, red, or purple, very often with callosities. Filaments subulate to linear; pollen 3-nucleate, exine tectum finely striate. Carpels connate for most of placental region, dehiscing distally in fruit; ovary subsuperior to subinferior. Integuments 2.

About 175 species: Asia, Europe, North America, mainly in arctic and montane regions; 166 species (112 endemic) in China.

Key 2

1 Leaf margin coarsely toothed or lobed.   (2)
+ Leaf margin entire.   (4)
2 (1) Leaf buds conspicuous in axils of bracts; sepals reflexed in fruit; petals white   108 S. strigosa
+ Leaf buds conspicuous or inconspicuous in axils of proximal leaves; sepals erect or spreading in fruit; petals yellow.   (3)
3 (2) Median leaves distributed evenly along stem, each usually with 3(or 5) apical lobes; inflorescence branches 1-flowered   106 S. hispidula
+ Median leaves aggregated, often into a rosette, each with several teeth; inflorescence branches 2- or more flowered   107 S. substrigosa
4 (1) Leaves pubescent adaxially.   (5)
+ Leaves glabrous adaxially.   (6)
5 (4) Leaf margin eglandular setose-ciliate; petals white   109 S. gemmipara
+ Leaf margin glandular hairy; petals yellow   105 S. balfourii
6 (4) Proximal leaves scalelike, with leaf buds in axils, median leaves best developed.   (7)
+ Proximal leaves nearly as well developed as median ones.   (10)
7 (6) Leaves linear, less than 1 mm wide.   (8)
+ Leaves ovate, oblanceolate, occasionally linear, but more than 1 mm wide.   (9)
8 (7) Stem many branched; leaf buds present in leaf and bract axils; pedicels glandular hairy; sepals glandular hairy abaxially; petals 2-4-callose   111 S. filicaulis
+ Stem simple; leaf buds absent in leaf and bract axils; pedicels glabrous; sepals glabrous on both surfaces; petals not callose   112 S. wenchuanensis
9 (7) Leaves oblong to narrowly subobovate, largest more than 8 mm, margin recurved   110 S. oreophila
+ Leaves suboblong or subspatulate, largest to 8 mm, margin flat   113 S. macrostigmatoides
10 (6) At least some leaves aggregated into an apparently basal rosette, distal cauline leaves ca. 1/2 size of rosette leaves   121 S. cinerascens
+ Leaves distributed ± evenly along stem, size differences gradual.   (11)
11 (10) Plants less than 3 cm tall; flower solitary, apparently sessile   120 S. erinacea
+ Plants more than 5 cm tall; flowers solitary or more than 1, pedicel at least 5 mm.   (12)
12 (11) Petal margin glandular ciliate or eglandular laciniate.   (13)
+ Petal margin entire, glabrous.   (14)
13 (12) Flower solitary   119 S. wardii
+ Flowers 2 or 3 in a compact corymb   118 S. gouldii
14 (12) Leaf margin not cartilaginous; leaf buds developing shoots more than 5 mm by anthesis   113 S. macrostigmatoides
+ Leaf margin cartilaginous; leaf buds ± undeveloped by anthesis.   (15)
15 (14) Broadest leaves at least 3 mm wide   114 S. wallichiana
+ Broadest leaves to 3 mm wide.   (16)
16 (15) Pedicels glabrous   116 S. brachypodoidea
+ Pedicels glandular hairy.   (17)
17 (16) Distal cauline leaves glandular hairy abaxially   117 S. zayuensis
+ Distal cauline leaves glabrous abaxially   115 S. brachypoda

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