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FOC | Family List | FOC Vol. 7 | Papaveraceae | Corydalis

31. Corydalis sect. Trachycarpae (Fedde) Fedde in Engler & Prantl, Nat. Pflanzenfam., ed. 2. 17b: 128. 1936.

糙果紫堇组 cao guo zi jin zu

Corydalis "Gruppe" Trachycarpae Fedde, Repert. Spec. Nov. Regni Veg. 20: 287. 1924; C. ser. Feddeanae C. Y. Wu & H. Chuang.

Herbs, perennial; stems and petioles of radical leaves attenuate to filiform underground base. Rhizome small, with fascicle of spindle-shaped to oblong, fleshy storage roots, apically with usually small pale overwintering bulblike bud. Stems simple or branched above. Radical leaves few. Cauline leaves 1-6. Leaflets often lanceolate, entire, with raised parallel veins abaxially. Fruiting pedicel recurved. Corolla usually yellow, rarely purple, very rarely blue, usually with distinct veins. Capsule explosively dehiscent.

Fifty-two species: a homogeneous and easily circumscribed section in C and W China, the Himalayas, and N India: 52 species (46 endemic) in China.


1 Fruiting pedicels erect in basal 2/3, curved at apex   (2)
+ Fruiting pedicels recurved from base   (3)
       
2 (1) Corolla pale to golden yellow; inner petals distinctly dark tipped; leaves 2-3 × pinnate.   238 C. scaberula
+ Corolla pale purple to purplish red, rarely pale yellow; inner petals not distinctly dark tipped; leaves 1-2 × pinnate.   237 C. glycyphyllos
       
3 (1) Flowers blue to purple   (4)
+ Flowers yellow   (21)
       
4 (3) Cauline leaves simply pinnate, or rarely innermost pinnae again divided; pinnae lanceolate-acute (cf. also   168 C. milarepa, roots unknown; stem leafy at base).
+ Cauline leaves bi- to triternate or bi- to tripinnate   (10)
       
5 Storage roots distinctly set off on long thin stalks   (6)
+ Storage roots sessile or with indistinct stalks   (8)
       
6 (5) Leaflets stiff, adaxially glossy dark green, abaxially with stiff 1-celled hairs on veins (Sichuan, NW Yunnan).   205 C. eugeniae
+ Leaflets slightly fleshy, glaucous, glabrous   (7)
       
7 (6) Bracts entire (N Sichuan).   207 C. degensis
+ Bracts divided (S Xizang).   196 C. sherriffii
       
8 (5) Spur broadly conical, 5-7 mm.   209 C. lathyrophylla
+ Spur longer   (9)
       
9 (8) Cauline leaves stiff, adaxially glossy, with narrow leaflets; bracts entire to divided.   203 C. atuntsuensis
+ Cauline leaves soft, glaucous, with elliptic leaflets; bracts entire, much shorter than pedicels.   197 C. kuruchuensis
       
10 (4) Racemes capitate; spur shorter than inner petals, downcurved.   208 C. brachyceras
+ Racemes not capitate or spur longer   (11)
       
11 (10) Capsule with lines of dense papillae   (12)
+ Capsule glabrous   (15)
       
12 (11) Corolla white, at least in early anthesis.   235 C. trachycarpa
+ Corolla purple to slate purplish blue   (13)
       
13 (12) Radical leaves bipinnate with 3 or 4 primary pinnae; upper petal 22-30 mm.   234 C. calcicola
+ Radical leaves bi- to tripinnate; upper petal shorter   (14)
       
14 (13) Leaves with 4-8 primary pinnae; upper petal 17-22 mm with spur 9-14 mm.   232 C. lhorongensis
+ Leaves with 3 or 4 pinnae; upper petal 12-13 mm with spur ca. 6 mm.   276 C. striatocarpa
       
15 (11) Cauline leaves stalked, bi- to triternate with stalked leaflets; stems slender   (16)
+ Cauline leaves bi- to tripinnate; stems slender or short   (17)
       
16 (15) Spur (4-)5-7 mm, narrow, straight.   188 C. lupinoides
+ Spur 16-18 mm, downcurved.   189 C. dorjii
       
17 (15) Stems 10-15 cm; plant of alpine scree   (18)
+ Stems 20-60 cm   (19)
       
18 (17) Spur ca. 18 mm; bracts entire.   233 C. dolichocentra
+ Spur 8-10 mm; bracts divided.   236 C. nubicola
       
19 (17) Spur 12-13 mm.   226 C. tsayulensis
+ Spur 14-17 mm   (20)
       
20 (19) Corolla light blue.   225 C. nemoralis
+ Corolla dark purple.   227 C. porphyrantha
       
21 (3) Cauline leaves simple   (22)
+ Cauline leaves compound   (23)
       
22 (21) Racemes 10-30-flowered; inner petals black tipped; bracts 5-20 mm.   191 C. juncea
+ Racemes 2-6-flowered; inner petals pale; bracts 15-50 mm.   192 C. pseudojuncea
       
23 (21) Storage roots with thin simple or branched stalks at least 10 mm (sometimes only stalks present in herbarium specimens)   (24)
+ Storage roots not or indistinctly stalked, or stalks shorter than 8 mm   (32)
       
24 (23) Cauline leaves pinnatifid, not cut quite to midvein, lower segments often again divided.   219 C. delavayi
+ Leaves pinnately to ternately divided with discrete leaflets   (25)
       
25 (24) Cauline leaves ternate to biternate or bipinnate; stem branched   (26)
+ Cauline leaves once pinnate; stem simple (rarely branched)   (28)
       
26 (25) Spur of upper petal 11-12 mm; plant 40-65 cm; racemes 9-25-flowered.   223 C. jiulongensis
+ Spur of upper petal 5-8 mm; plant 10-35 cm; racemes 2-9-flowered   (27)
       
27 (26) Cauline leaves stalked; leaflets divided into obovate to oblanceolate lobes.   224 C. homopetala
+ Cauline leaves sessile; leaflets linear to narrowly lanceolate.   193 C. lowndesii
       
28 (25) Raceme 10-25-flowered; spur 6-9 mm   (29)
+ Raceme 4-10-flowered   (30)
       
29 (28) Spur upwardly directed, broadly conical.   206 C. prattii
+ Spur of upper petal horizontal.   194 C. polygalina
       
30 (28) Inner petals 6-7 mm; spur 6-9 mm.   194 C. polygalina
+ Inner petals 8-10 mm; spur 7-12 mm   (31)
       
31 (30) Bracts entire.   204 C. mayae
+ Lower bracts divided.   205 C. eugeniae
       
32 (23) Most bracts divided   (33)
+ Bracts entire or lowermost sometimes divided; stem usually simple (but sometimes branched in C. kingdonis, C. quinquefoliolata, C. sophronitis, and C. trisecta)   (44)
       
33 (32) Cauline leaves simply pinnate with narrow entire leaflets; stems simple   (34)
+ Cauline leaves more divided; stem branched, rarely simple   (36)
       
34 (33) Crest of outer petals broad, rounded, much overtopping apex; spur strongly sigmoidally curved, 12-15 mm.   199 C. rheinbabeniana
+ Crest of outer petals not overtopping apex; spur straight to slightly curved, 8-11 mm   (35)
       
35 (34) Leaflets glossy green adaxially, stiff; storage roots oblong.   203 C. atuntsuensis
+ Leaflets glaucous, soft; storage roots fusiform.   202 C. linarioides
       
36 (33) Ovary and fruit with lines of papillae   (37)
+ Fruit glabrous   (39)
       
37 (36) Spur of upper petal 15-18 mm, tapering to narrow apex.   231 C. chamdoensis
+ Spur shorter   (38)
       
38 (37) Spur of upper petal 12-14 mm.   230 C. temolana
+ Spur of upper petal 8-10 mm.   229 C. stramineoides
       
39 (36) Racemes ± corymbose; lower pedicels 10-25 mm; stems 5-15 cm above soil surface   (40)
+ Racemes elongate; lower pedicels 5-12 mm; stems 15-65 cm above soil surface   (42)
       
40 (39) Upper petal entire; crest attenuate to both ends; spur tapering to acute apex; plant glabrous.   220 C. tenuipes
+ Upper petal dentate; crest broadly obtuse, ± dentate; spur cylindric; plant usually papillose-hairy   (41)
       
41 (40) Spur 10-11 mm, slightly upturned.   239 C. petrodoxa
+ Spur 6-9 mm, downcurved at apex.   221 C. oligosperma
       
42 (39) Racemes lax, 5-7-flowered; spur 7-8 mm.   224 C. homopetala
+ Racemes dense, 10-30-flowered; spur longer   (43)
       
43 (42) Spur 14-16 mm, cylindric, usually strongly recurved; stems usually simple, erect; corolla pale yellow.   222 C. pseudodensispica
+ Spur tapering to apex, 8-13 mm; stems branched, ascending to suberect; corolla bright yellow, with darker veins.   228 C. densispica
       
44 (32) Spur of upper petal 5-8 mm   (45)
+ Spur of upper petal 9-20 mm   (53)
       
45 (44) Inner petals tipped with contrasting dark purple   (46)
+ Inner petals pale at apex   (47)
       
46 (45) Cauline leaves pinnate, sessile; leaflets linear.   210 C. helodes
+ Cauline leaves biternate, shortly stalked; leaflets lanceolate.   190 C. kingdonis
       
47 (45) Crest of upper petal broad, clearly overtopping apex   (48)
+ Crest not overtopping apex, or absent   (50)
       
48 (47) Lower petal conspicuously saccate; bracts sharply reflexed in fruit.   215 C. gaoxinfeniae
+ Lower petal not or slightly saccate; bracts not sharply reflexed in fruit   (49)
       
49 (48) Spur broad, cylindric, straight; stems simple.   214 C. linearis
+ Spur slender, tapering, curved; stem ± branched.   200 C. sophronitis
       
50 (47) Upper petal ca. 13 mm   (51)
+ Upper petal 15-17 mm   (52)
       
51 (50) Outer petals without crest; stigma square.   218 C. mediterranea
+ Outer petals crested; stigma much broader than long with distinct marginal papillae, geminate papillae lacking or indistinct.   195 C. quinquefoliolata
       
52 (50) Leaflets linear to narrowly lanceolate, stiff, glossy green adaxially.   203 C. atuntsuensis
+ Leaflets broadly elliptic, fleshy, glaucous.   198 C. trisecta
       
53 (44) Lower petal not or very slightly saccate   (54)
+ Lower petal broadly saccate   (57)
       
54 (53) Leaflets glossy green adaxially, stiff; storage roots oblong.   203 C. atuntsuensis
+ Leaflets glaucous, soft; storage roots fusiform   (55)
       
55 (54) Spur of upper petal tapering to apex, 8-10 mm.   198 C. trisecta
+ Spur of upper petal cylindric   (56)
       
56 (55) Stem 8-20 cm; bracts obovate, 2-9 mm wide; upper petal 22-27 mm.   201 C. cristata
+ Stem 10-50 cm; bracts linear-lanceolate, 1-3 mm wide; upper petal 16-23 mm.   202 C. linarioides
       
57 (53) Bracts sharply reflexed in fruit; spur slightly to strongly downcurved   (58)
+ Bracts not sharply reflexed in fruit; spur sigmoidally curved to straight   (59)
       
58 (57) Spur slightly downcurved, tapering, 11-15 mm; leaflets of cauline leaves 2-5 mm wide.   216 C. feddeana
+ Spur strongly curved, cylindric, 15-20 mm; leaflets 3-14 mm wide.   217 C. pseudocristata
       
59 (57) Inner petals pale; leaflets linear, 20-50 mm.   213 C. sigmoides
+ Inner petals with contrasting dark purple apex; ultimate leaflets shorter   (60)
       
60 (59) Spur attenuate to thin apex; cauline leaves biternate.   211 C. lopinensis
+ Spur cylindric or slightly attenuate; cauline leaves tripinnate.   212 C. bijiangensis

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