| 1 |
Phyllary margins pectinate, with 2-5 pairs of teeth |
|
(2) |
| + |
Phyllary margins entire or subentire |
|
(3) |
| |
|
|
|
| 2 (1) |
Involucre 5-8 mm in diam.; phyllaries with 2 or 3 pairs of teeth; leaves with 8-17 pairs of lobes, lobes oblong to narrowly elliptic-linear. |
|
207 S. odontolepis |
| + |
Involucre 8-12 mm in diam.; phyllaries with 3-5 pairs of teeth; leaves with 4-9 pairs of lobes, lobes ovate to narrowly ovate-elliptic. |
|
208 S. pectinata |
| |
|
|
|
| 3 (1) |
Basal and lower stem leaves with a shallowly or deeply cordate base |
|
(4) |
| + |
Basal and lower leaves with an attenuate, cuneate, rounded, truncate, or auriculate-amplexicaul base |
|
(30) |
| |
|
|
|
| 4 (3) |
Leaf blade adaxially green, abaxially grayish white or grayish green and densely tomentose, arachnoid, or lanate |
|
(5) |
| + |
Leaf blade green, concolorous or abaxially paler green, abaxially glabrous or glabrescent |
|
(13) |
| |
|
|
|
| 5 (4) |
Basal and lower stem leaves narrowly ovate-elliptic, 0.7-2.5 cm wide. |
|
226 S. graciliformis |
| + |
Basal and lower stem leaves 2-17 cm wide |
|
(6) |
| |
|
|
|
| 6 (5) |
Lower stem leaves triangular-ovate, base cordate or hastate |
|
(7) |
| + |
Lower stem leaves ovate, ovate-elliptic, or reniform |
|
(8) |
| |
|
|
|
| 7 (6) |
Leaf blade leathery, adaxially pale green or grayish green, sparsely arachnoid floccose, glabrescent; involucre narrowly campanulate, 4-7 mm in diam. |
|
225 S. firma |
| + |
Leaf blade papery, adaxially white or grayish white, densely tomentose; involucre campanulate, 6-12 mm in diam. |
|
227 S. nivea |
| |
|
|
|
| 8 (6) |
Apex of phyllaries without a mucro |
|
(9) |
| + |
Apex of phyllaries with a black mucro |
|
(10) |
| |
|
|
|
| 9 (8) |
Stem much branched; capitula in paniculiform synflorescence; involucre cylindric, 0.7-0.9 cm in diam. |
|
233 S. pinetorum |
| + |
Stem apically few branched; capitula in racemiform or lax corymbiform synflorescence; involucre campanulate-obconic, 1.2-1.5 cm in diam. |
|
235 S. chetchozensis |
| |
|
|
|
| 10 (8) |
Stem densely leafy; capitula numerous, in paniculiform to lax corymbiform synflorescence. |
|
229 S. vestitiformis |
| + |
Stem with few leaves; capitula few or numerous, in clustered corymbiform synflorescence |
|
(11) |
| |
|
|
|
| 11 (10) |
Stem with brown multicellular articulate hairs. |
|
230 S. baroniana |
| + |
Stem white arachnoid or lanate |
|
(12) |
| |
|
|
|
| 12 (11) |
Leaf blade 2.5-6 × 2-5 cm; stem slender; involucre obconic to cylindric, 5-6 mm in diam. |
|
231 S. flaccida |
| + |
Leaf blade 3-11 × 7-10(-17) cm; stem stout; involucre campanulate, 6-10 mm in diam. |
|
232 S. acromelaena |
| |
|
|
|
| 13 (4) |
Capitulum usually solitary, terminal on stem, sometimes 2-5, solitary on end of branches; involucre 2-2.5 cm in diam. |
|
209 S. populifolia |
| + |
Capitula usually numerous or few, rarely solitary; involucre usually less than 2 cm in diam |
|
(14) |
| |
|
|
|
| 14 (13) |
Apical part of phyllaries either herbaceous or leathery and saber-shaped, usually patent or reflexed |
|
(15) |
| + |
Apical part of phyllaries neither herbaceous nor leathery and saber-shaped, erect, usually appressed |
|
(21) |
| |
|
|
|
| 15 (14) |
Apical part of phyllaries leathery, saber-shaped and reflexed, apex long acuminate |
|
(16) |
| + |
Apical part of phyllaries herbaceous, not saber-shaped, reflexed, patent, or erect, apex shortly acuminate |
|
(18) |
| |
|
|
|
| 16 (15) |
Leaf blade undivided but margin dentate. |
|
212 S. subtriangulata |
| + |
Leaf blade pinnately lobed or pinnately dentate |
|
(17) |
| |
|
|
|
| 17 (16) |
Capitula numerous, shortly pedunculate; involucre narrowly campanulate, 5-7 mm in diam. |
|
210 S. mongolica |
| + |
Capitula few, long pedunculate; involucre obconic, 10-15 mm in diam. |
|
211 S. sinuata |
| |
|
|
|
| 18 (15) |
Leaf blade hastate or narrowly cordate, longer than wide |
|
(19) |
| + |
Leaf blade cordate or broadly cordate, ± as long as wide |
|
(20) |
| |
|
|
|
| 19 (18) |
Capitula 6-12, in paniculiform to corymbiform synflorescence; involucre narrowly campanulate, 0.7-1 cm in diam. |
|
215 S. stricta |
| + |
Capitula 1-3, long pedunculate, in corymbiform synflorescence; involucre campanulate, 1-2 cm in diam. |
|
216 S. hylophila |
| |
|
|
|
| 20 (18) |
Lower and middle stem leaves broadly cordate, 8-20 × 7-18 cm; stem glabrous. |
|
213 S. cordifolia |
| + |
Lower and middle stem leaves ovate-cordate, 5-11 × 5-11 cm; stem with sparse articulate hairs but glabrescent. |
|
214 S. oligantha |
| |
|
|
|
| 21 (14) |
Involucre ca. 5 mm in diam.; corolla ca. 8 mm; inner pappus bristles 6-7 mm. |
|
221 S. tenerifolia |
| + |
Involucre 5-15 mm in diam.; corolla 10-16 mm; inner pappus bristles 7-13 mm |
|
(22) |
| |
|
|
|
| 22 (21) |
Involucre campanulate to globose, 10-15 mm in diam |
|
(23) |
| + |
Involucre tubular to campanulate, 5-10 mm in diam |
|
(25) |
| |
|
|
|
| 23 (22) |
Phyllaries long acuminate. |
|
223 S. hwangshanensis |
| + |
Phyllaries acute to obtuse and sometimes mucronate |
|
(24) |
| |
|
|
|
| 24 (23) |
Leaf blade sparsely strigose on both surfaces. |
|
222 S. grandifolia |
| + |
Leaf blade abaxially thinly arachnoid tomentose but glabrescent, adaxially sparsely strigose. |
|
219 S. bullockii |
| |
|
|
|
| 25 (22) |
Leaf blade leathery, abaxially grayish green and sparsely arachnoid floccose but glabrescent. |
|
225 S. firma |
| + |
Leaf blade papery, abaxially pale green and subglabrous |
|
(26) |
| |
|
|
|
| 26 (25) |
Involucre campanulate, 8-10 mm in diam |
|
(27) |
| + |
Involucre tubular to narrowly campanulate, 5-8 mm in diam |
|
(28) |
| |
|
|
|
| 27 (26) |
Phyllaries appressed, not patent or recurved. |
|
220 S. pseudobullockii |
| + |
Phyllaries apically patent to recurved. |
|
234 S. paleata |
| |
|
|
|
| 28 (26) |
Plants 30-40 cm tall; stem simple or apically few branched; capitula 3-5, subsessile in a clustered corymbiform synflorescence. |
|
218 S. triangulata |
| + |
Plants 50-120 cm tall; stem much branched; capitula more numerous, shortly or long pedunculate |
|
(29) |
| |
|
|
|
| 29 (28) |
Capitula shortly pedunculate, clustered at end of branches in a corymbiform synflorescence. |
|
224 S. ussuriensis |
| + |
Capitula long pedunculate, in a paniculiform synflorescence. |
|
217 S. manshurica |
| |
|
|
|
| 30 (3) |
Leaf blade base auriculate and amplexicaul |
|
(31) |
| + |
Leaf blade base not auriculate or amplexicaul |
|
(37) |
| |
|
|
|
| 31 (30) |
Capitula ca. 5, sessile, clustered in a hemispheric synflorescence. |
|
236 S. lyratifolia |
| + |
Capitula either few and long pedunculate or numerous and shortly pedunculate, in a corymbiform synflorescence |
|
(32) |
| |
|
|
|
| 32 (31) |
Capitula 1-4, large, long pedunculate, not in a clustered corymbiform synflorescence; involucre 1.2-2 cm in diam |
|
(33) |
| + |
Capitula numerous, small or medium-sized, shortly pedunculate, in a clustered corymbiform synflorescence; involucre 0.4-1 cm in diam |
|
(34) |
| |
|
|
|
| 33 (32) |
Leaf blade ovate to narrowly ovate-elliptic, 1.5-5 cm wide. |
|
237 S. neofranchetii |
| + |
Leaf blade narrowly oblong to linear, 1-1.5 cm wide. |
|
238 S. caudata |
| |
|
|
|
| 34 (32) |
Leaf blade base inconspicuously auriculate |
|
(35) |
| + |
Leaf blade base broadly auriculate and amplexicaul |
|
(36) |
| |
|
|
|
| 35 (34) |
Stem winged; leaf blade abaxially grayish green or grayish white, densely arachnoid tomentose. |
|
239 S. leclerei |
| + |
Stem wingless; leaf blade abaxially green, puberulent only on veins. |
|
240 S. merinoi |
| |
|
|
|
| 36 (34) |
All leaves sessile; leaf blade elliptic, 8-23 × 2.5-6 cm. |
|
241 S. macrota |
| + |
Lower stem leaves petiolate; leaf blade narrowly ovate-elliptic, 6-9 × 2.5-4 cm. |
|
242 S. semiamplexicaulis |
| |
|
|
|
| 37 (30) |
Leaf blade lyrate or lyrate pinnatilobed; petiole with 1 or several pairs of lobes |
|
(38) |
| + |
Leaf blade undivided or pinnately lobed but not lyrate; petiole without lobes |
|
(50) |
| |
|
|
|
| 38 (37) |
Receptacle without bristles. |
|
178 S. porphyroleuca (see S. sect. Strictae) |
| + |
Receptacle with bristles |
|
(39) |
| |
|
|
|
| 39 (38) |
Leaf blade decurrent on stem; stem winged |
|
(40) |
| + |
Leaf blade not decurrent on stem; stem wingless |
|
(45) |
| |
|
|
|
| 40 (39) |
Leaf blade glabrous or subglabrous on both surfaces |
|
(41) |
| + |
Leaf blade adaxially green and glabrous or pubescent, abaxially grayish white and arachnoid or lanate |
|
(42) |
| |
|
|
|
| 41 (40) |
Outer phyllaries acute to obtuse. |
|
243 S. peduncularis |
| + |
Outer phyllaries apically with a 3-4 mm subulate awn. |
|
244 S. frondosa |
| |
|
|
|
| 42 (40) |
Involucre campanulate, 1-1.7 cm in diam. |
|
245 S. eriocephala |
| + |
Involucre ovoid or campanulate, 0.5-0.8 cm in diam |
|
(43) |
| |
|
|
|
| 43 (42) |
Plants 5-15 cm tall. |
|
261 S. kiraisiensis |
| + |
Plants 20-90 cm tall |
|
(44) |
| |
|
|
|
| 44 (43) |
Stem wings entire; leaf blade ovate, undivided. |
|
228 S. dimorphaea |
| + |
Stem wings sinuate-dentate; leaf blade narrowly ovate-elliptic, runcinate-pinnately lobed. |
|
246 S. dzeurensis |
| |
|
|
|
| 45 (39) |
Leaf blade adaxially green and glabrous, abaxially grayish white or grayish green and arachnoid tomentose |
|
(46) |
| + |
Leaf blade green, ± concolorous, glabrous or subglabrous |
|
(48) |
| |
|
|
|
| 46 (45) |
Terminal lobe of leaf blade ovate, base cuneate to subtruncate. |
|
131 S. flexuosa (see S. sect. Laguranthera) |
| + |
Terminal lobe of leaf blade ovate-triangular or ovate, base conspicuously cordate |
|
(47) |
| |
|
|
|
| 47 (46) |
Stem with few reduced leaves; basal leaves numerous, in a basal rosette. |
|
198 S. alatipes (see S. sect. Rosulascentes) |
| + |
Stem with many well-developed leaves; basal leaves few, usually withered at anthesis. |
|
250 S. lampsanifolia |
| |
|
|
|
| 48 (45) |
Capitulum solitary, terminal on stem, or 2-4; phyllaries long acuminate, reflexed. |
|
249 S. morifolia |
| + |
Capitula numerous, in a racemiform or corymbiform synflorescence; phyllaries not reflexed |
|
(49) |
| |
|
|
|
| 49 (48) |
Involucre hemispheric, 2-2.5 cm in diam. |
|
247 S. carduiformis |
| + |
Involucre narrowly campanulate, 0.4-0.9 cm in diam. |
|
248 S. mutabilis |
| |
|
|
|
| 50 (37) |
Leaf blade pinnately lobed or coarsely pinnately dentate |
|
(51) |
| + |
Leaf blade undivided, margin entire, denticulate, or shallowly sinuate-dentate |
|
(52) |
| |
|
|
|
| 51 (50) |
Plants 15-35 cm tall; capitulum solitary. |
|
286 S. grosseserrata |
| + |
Plants 40-100 cm tall; capitula 3-5 in a clustered corymbiform synflorescence. |
|
289 S. recurvata |
| |
|
|
|
| 52 (50) |
Leaf blade adaxially green, smooth, and glabrous or scabrous and strigose, abaxially white, grayish white, or grayish green and densely tomentose or thinly arachnoid |
|
(53) |
| + |
Leaf blade green, concolorous, glabrous, sparsely pubescent, or glandular hairy |
|
(69) |
| |
|
|
|
| 53 (52) |
Plants stemless or shortly stemmed, 3-8 cm tall; capitulum solitary; involucre 2-2.5 cm in diam. |
|
272 S. woodiana |
| + |
Plants always with a developed stems, more than 10 cm tall; capitula usually several; involucre less than 1.5 cm in diam |
|
(54) |
| |
|
|
|
| 54 (53) |
Capitula 1-6 |
|
(55) |
| + |
Capitula numerous, in a racemiform, paniculiform, or corymbiform synflorescence |
|
(59) |
| |
|
|
|
| 55 (54) |
Stem and leaf petioles wingless |
|
(56) |
| + |
Stem and leaf petioles distinctly winged |
|
(57) |
| |
|
|
|
| 56 (55) |
Plants 40-65 cm tall; involucre 0.8-1 cm in diam. |
|
270 S. iodoleuca |
| + |
Plants 10-30 cm tall; involucre 1-1.2 cm in diam. |
|
275 S. alaschanica |
| |
|
|
|
| 57 (55) |
Plants 30-40 cm tall; leaf blade ovate to elliptic. |
|
271 S. smithiana |
| + |
Plants 10-35 cm tall; leaf blade broadly ovate to orbicular |
|
(58) |
| |
|
|
|
| 58 (57) |
Involucre narrowly campanulate to cylindric, 5-7 mm in diam. |
|
273 S. komarnitzkii |
| + |
Involucre ovoid to obconic, ca. 13 mm in diam. |
|
274 S. rotundifolia |
| |
|
|
|
| 59 (54) |
Stem wingless |
|
(60) |
| + |
Stem winged |
|
(64) |
| |
|
|
|
| 60 (59) |
Leaf blade orbicular to broadly ovate, abaxially white and densely tomentose. |
|
281 S. bullata |
| + |
Leaf blade ovate, elliptic, or oblong to narrowly elliptic, abaxially grayish green and arachnoid |
|
(61) |
| |
|
|
|
| 61 (60) |
Plants to 1.2 m tall; capitula in a paniculiform to lax corymbiform synflorescence. |
|
285 S. undulata |
| + |
Plants to 70 cm tall; capitula in a clustered corymbiform synflorescence |
|
(62) |
| |
|
|
|
| 62 (61) |
Leaf blade ovate to elliptic, adaxially grayish green, and arachnoid floccose. |
|
283 S. elliptica |
| + |
Leaf blade narrowly ovate-elliptic to narrowly elliptic, adaxially green and glabrous |
|
(63) |
| |
|
|
|
| 63 (62) |
Plants 30-70 cm tall; leaves and phyllaries usually without glands. |
|
282 S. alpina |
| + |
Plants 10-20 cm tall; leaves and phyllaries abaxially densely gland-dotted. |
|
284 S. pseudoalpina |
| |
|
|
|
| 64 (59) |
Capitula 4-10, in a lax racemiform synflorescence. |
|
276 S. polygonifolia |
| + |
Capitula usually more numerous, in a clustered or lax corymbiform or paniculiform synflorescence |
|
(65) |
| |
|
|
|
| 65 (64) |
Stem stout, much branched; leaf blade oblong to narrowly elliptic, less than 2 cm wide. |
|
278 S. platypoda |
| + |
Stem slender, simple or apically few branched; leaf blade ovate to narrowly elliptic, usually more than 2 cm wide |
|
(66) |
| |
|
|
|
| 66 (65) |
Middle stem leaves 2-4 cm wide, apex acuminate |
|
(67) |
| + |
Middle stem leaves 3-7.5 cm wide, apex acute to obtuse |
|
(68) |
| |
|
|
|
| 67 (66) |
Middle stem leaves narrowly elliptic to linear, 10-20 cm. |
|
280 S. amurensis |
| + |
Middle stem leaves ovate to elliptic, 5-8 cm. |
|
228 S. dimorphaea |
| |
|
|
|
| 68 (66) |
Phyllaries yellowish green, apically purplish, densely arachnoid pubescent; outer phyllaries acuminate but without a black mucro; leaf blade apex obtuse to subacute. |
|
277 S. vestita |
| + |
Phyllaries straw-colored, brownish, or purple, pubescent or arachnoid; outer phyllaries with a long black mucro; leaf blade apex acute. |
|
279 S. cauloptera |
| |
|
|
|
| 69 (52) |
Capitulum solitary, terminal on stem |
|
(70) |
| + |
Capitula 2-6 or numerous, in a corymbiform synflorescence |
|
(77) |
| |
|
|
|
| 70 (69) |
Leaf blade margin coarsely sinuate-dentate |
|
(71) |
| + |
Leaf blade margin entire or inconspicuously denticulate |
|
(72) |
| |
|
|
|
| 71 (70) |
Phyllaries not reflexed; leaf blade obovate-elliptic, apex rounded to acute and mucronate. |
|
286 S. grosseserrata |
| + |
Phyllaries reflexed; leaf blade ovate, narrowly ovate-elliptic, or triangular, apex acuminate. |
|
288 S. tunglingensis |
| |
|
|
|
| 72 (70) |
Plants stemless or with a developed stem; involucre 1.5-4 cm in diam. |
|
252 S. superba |
| + |
Plants always with a developed stem; involucre 1-2 cm in diam |
|
(73) |
| |
|
|
|
| 73 (72) |
Leaf blade oblong to broadly linear, margin sinuate. |
|
287 S. pratensis |
| + |
Leaf blade narrowly obovate, narrowly ovate, or narrowly elliptic, margin entire or denticulate |
|
(74) |
| |
|
|
|
| 74 (73) |
Leaf blade narrowly oblong to linear, glabrous; phyllaries reflexed. |
|
257 S. micradenia |
| + |
Leaf blade narrowly ovate-elliptic to narrowly elliptic, sparsely villous; phyllaries not reflexed |
|
(75) |
| |
|
|
|
| 75 (74) |
Involucre obconic; phyllaries glabrous; leaf blade margin denticulate. |
|
254 S. acutisquama |
| + |
Involucre campanulate; phyllaries villous; leaf blade margin entire and densely ciliate |
|
(76) |
| |
|
|
|
| 76 (75) |
Leaf blade base oblique and rounded or cuneate, apex obtuse to acute; phyllaries densely villous; corolla 1.8-2.4 cm. |
|
251 S. hieracioides |
| + |
Leaf blade base attenuate, apex acute to acuminate; phyllaries sparsely villous; corolla 1.2-1.4 cm. |
|
253 S. tatsienensis |
| |
|
|
|
| 77 (69) |
Capitula 2-6 |
|
(78) |
| + |
Capitula numerous |
|
(83) |
| |
|
|
|
| 78 (77) |
Leaf blade margin retrorsely dentate or serrate. |
|
258 S. retroserrata |
| + |
Leaf blade margin entire or dentate to serrate but not retrorsely so |
|
(79) |
| |
|
|
|
| 79 (78) |
Leaf blade broadly ovate to narrowly elliptic, more than 3 cm wide, margin dentate to serrate |
|
(80) |
| + |
Leaf blade narrowly ovate, narrowly elliptic, narrowly oblong, or linear, 0.8-2.5 cm wide, margin entire or denticulate |
|
(81) |
| |
|
|
|
| 80 (79) |
Middle and upper stem leaves narrowly ovate to narrowly elliptic. |
|
256 S. hultenii |
| + |
Middle and upper stem leaves broadly ovate to ovate-elliptic. |
|
255 S. euodonta |
| |
|
|
|
| 81 (79) |
Leaf blade narrowly oblong to linear, glabrous; phyllaries reflexed. |
|
257 S. micradenia |
| + |
Leaf blade narrowly ovate-elliptic to narrowly elliptic, sparsely villous; phyllaries not reflexed |
|
(82) |
| |
|
|
|
| 82 (81) |
Involucre campanulate; phyllaries villous; leaf blade margin entire and densely ciliate. |
|
253 S. tatsienensis |
| + |
Involucre obconic; phyllaries glabrous; leaf blade margin denticulate. |
|
254 S. acutisquama |
| |
|
|
|
| 83 (77) |
Phyllaries apically saber-shaped |
|
(84) |
| + |
Phyllaries apically not saber-shaped |
|
(86) |
| |
|
|
|
| 84 (83) |
Stem and petioles wingless. |
|
262 S. epilobioides |
| + |
Stem and petioles narrowly winged |
|
(85) |
| |
|
|
|
| 85 (84) |
Leaf blade margin denticulate; involucre 5-6 mm in diam. |
|
259 S. acuminata |
| + |
Leaf blade margin dentate; involucre ca. 10 mm in diam. |
|
263 S. umbrosa |
| |
|
|
|
| 86 (83) |
Stem and petioles wingless; leaf blade triangular-ovate to ovate. |
|
268 S. dielsiana |
| + |
Stem and petioles distinctly winged; leaf blade obovate to elliptic or narrowly so |
|
(87) |
| |
|
|
|
| 87 (86) |
Leaf blade abaxially glandular; involucre ca. 3 mm in diam.; phyllaries acuminate. |
|
260 S. glandulosa |
| + |
Leaf blade abaxially not glandular; involucre 5-8 mm in diam.; phyllaries acute to obtuse |
|
(88) |
| |
|
|
|
| 88 (87) |
Leaf blade 3-9 cm wide |
|
(89) |
| + |
Leaf blade 1-4 cm wide |
|
(90) |
| |
|
|
|
| 89 (88) |
Leaf blade green, concolorous; lower stem leaves with base cordate and apex acute to acuminate. |
|
267 S. latifolia |
| + |
Leaf blade adaxially green, abaxially glaucous-green; lower stem leaves with base attenuate and apex very long acuminate. |
|
269 S. neoserrata |
| |
|
|
|
| 90 (88) |
Outer phyllaries with apex acuminate. |
|
266 S. candolleana |
| + |
Outer phyllaries with apex obtuse or subacute |
|
(91) |
| |
|
|
|
| 91 (90) |
Leaf blade of middle stem leaves ovate, base abruptly narrowed into a winged petiole to 5.5 cm; leaf blade adaxially smooth; phyllaries usually yellowish green. |
|
264 S. hemsleyi |
| + |
Leaf blade of middle stem leaves narrowly elliptic, base attenuate into a petiole to 1(-2) cm; leaf blade adaxially scabrid; phyllaries usually dark purple. |
|
265 S. parviflora |