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FOC | Family List | FOC Vol. 20-21 | Asteraceae

151. Artemisia Linnaeus, Sp. Pl. 2: 845. 1753.

蒿属 hao shu

Authors: Yourun Lin(Ling Yuou-ruen), Christopher J. Humphries & Michael G. Gilbert

Herbs, annual or perennial, subshrubs, or shrubs, usually strongly and pleasantly aromatic, indumentum absent or of basifixed, medifixed, gland-tipped or viscid hairs. Leaves alternate, pinnate, rarely palmately divided or entire. Synflorescences racemose, sometimes spicate, usually grouped into panicles; capitula usually many, often secund, usually small, shortly pedunculate to sessile, heterogamous, disciform. Involucres globose, ovoid, or ellipsoid; phyllaries in 3 or 4 rows, completely scarious or herbaceous with broad to narrow scarious margin. Receptacle convex or flat, epaleate, glabrous or pubescent. Marginal florets in 1(or 2) series, 3-10 or more, female; corolla tubular, rarely vasiform, cup-shaped, or conical, apex 2(-4)-toothed; style exserted, apex acute. Disk florets several to many, in 2 or more series, male or bisexual; corolla tubular, apex 5-toothed. Anthers with 2 obtuse basal appendages, apical appendage acute, triangular. Style ca. as long as or longer than corolla, divergent, and with a truncate or folding and pedunculate apex, or sometimes shorter than corolla, not divergent, apex clavate or funnelform. Achenes obovoid, ovoid, or oblong, faintly striate. Corona absent or minute.

About 380 species: mainly in the N Hemisphere, also in Africa, Australia, and Central and South America; 186 species (82 endemic) in China.

Many species are rich in polyacetylenes, flavonoids, terpenoids, and cyanogenic glycosides and are well-known medicinal plants. Drugs based on artemisinin, originally derived from Artemisia annua, are particularly important for the treatment of chloroquine-resistant strains of malaria. Artemisia abrotanum Linnaeus, A. absinthium, A. dracunculus, and A. vulgaris are widely cultivated as culinary and medicinal herbs. Some are used to stabilize sand in desert or semidesert areas or as herbage for feeding livestock.

A possible record of Artemisia quinqueloba Trautvetter from China (Pampanini, Nuovo Giorn. Bot. Ital., n.s., 34: 685. 1927) cannot be confirmed, as the specimen on which it was based, "Songoria, Schrenk" (B), is no longer extant.


1 Disk florets male, ovaries usually minute or absent; style shorter than corolla, usually not divergent, 2-cleft, with clavate or funnelform apex   (2)
+ Disk florets bisexual, ovaries well developed; style ca. as long as or longer than corolla, branches divergent, recurved at apex   (3)
       
2 (1) Lobules of leaf blade filiform, narrowly linear, lanceolate, or subulate and less than 1.5 mm wide, or pectinate and 1-2.5 × 1-2.5 mm, or leaf blade lanceolate or linear-lanceolate; ovaries of disk florets usually minute.   6 A. sect. Dracunculus (p. 719)
+ Lobules of leaf blade broadly linear to lanceolate or elliptic, more than (1-)1.5 mm wide, or leaf blade spatulate or obovate; ovaries of disk florets usually absent.   7 A. sect. Latilobus (p. 731)
       
3 (1) Receptacle pubescent or chaffy, sometimes hairs deciduous; corolla of female florets usually vasiform or narrowly conical, (2-)4-toothed.   1 A. sect. Absinthium (p. 677)
+ Receptacle glabrous; corolla of female florets narrowly tubular or rarely narrowly conical, 2- or 3(or 4)-toothed or without teeth   (4)
       
4 (3) Phyllaries scarious throughout, glabrescent, midvein not green.   5 A. sect. Albibractea (p. 717)
+ Phyllaries scarious only at margin, midvein green or sometimes brown   (5)
       
5 (4) Plants glandular or viscid pubescent.   4 A. sect. Viscidipubes (p. 711)
+ Plants not glandular or viscid, puberulent or glabrescent   (6)
       
6 (5) Capitula globose, rarely ovoid; lobules of leaf blade less than 1(-1.5) mm wide, if wider then pectinate.   2 A. sect. Abrotanum (p. 685)
+ Capitula ellipsoid or oblong, rarely subglobose, campanulate, or hemispheric; lobules of leaf blade more than (1.5-)2 mm wide.   3 A. sect. Artemisia (p. 694)

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