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Lamiaceae Lindley

唇形科 chun xing ke

Authors: Xi-wen Li & Ian C. Hedge

Salvia przewalskii

Credit: Harvard University Herbaria

Herbs, sometimes subshrubs or shrubs, annual or perennial, usually aromatic. Stems and branches usually 4-angled. Leaves opposite, rarely whorled or alternate, simple to pinnately dissected or compound, without stipules. Inflorescences generally compound, sometimes flowers solitary and axillary; verticillasters 2- to many flowered, subtended by leaves or bracts. Flowers bisexual, zygomorphic, rarely subactinomorphic, bracteolate or not. Calyx persistent, 5-toothed, 2-lipped; upper lip 3-toothed or entire (deciduous in Scutellaria); lower lip 2- or 4-toothed; tube sometimes hairy annulate inside. Corolla limb usually 2-lipped; upper lip 2-lobed and lower 3-lobed, rarely upper lip entire and lower 4-lobed, also rarely limb (4- or) 5-lobed; tube hairy annulate inside. Stamens epipetalous, 4 or 2, free, rarely filaments connate, sometimes one staminodial; anther 1- or 2-celled, usually dehiscing longitudinally; disc persistent. Ovary superior, 2-celled and each cell 2-ovuled and style subterminal, or ovary 4-parted and each lobe 1-ovuled and style gynobasic (from bases of ovary lobes) with 2-cleft apex. Fruit usually 4 dry nutlets. Seeds with or without endosperm.

Approximately 3500 species in 220 genera, distributed worldwide, but mostly in the Mediterranean region and SW Asia. China has 807 species in 96 genera.

Cantino (Taxon 40: 441. 1991) suggests that Cardioteucris cordifolia C. Y. Wu, originally placed in the Lamiaceae, is identical with Caryopteris siccanea W. Smith (Verbenaceae, q.v.). Although the original author agrees with the identity of Cardioteucris cordifolia and Cardiopteris siccaneae, he believes that the generic placement of Cardioteucris is in the Lamiaceae because of its 2-lipped calyx and deeply 4-divided ovary. The last feature is aberrant in Caryopteris and all other Verbenaceae.

Wu Cheng-yih & Li Hsi-wen, eds. 1977. Labiatae. Fl. Reipubl. Popularis Sin. 65(2): 1-649, 66: 1-647.


Key 3

1 Corolla resupinate; calyx tube hairy annulate at middle or on throat inside.   (2)
+ Corolla not resupinate; calyx tube not hairy annulate at middle inside.   (3)
       
2 (1) Calyx with 5 subequal teeth; erect herbs; leaves narrowly to broadly ovate differentiated in to stem and floralones   17 Lophanthus
+ Calyx 2-lipped, upper lip 3-toothed, lower lip 2-toothed; stem prostrate, basally leafless, leaves dense and decussate apically   21 Marmoritis
       
3 (1) Stamen pairs not parallel to each other, posture various but not all arcuate ascending under upper lip of corolla   16 Agastache
+ Stamen pairs parallel to each other, all arcuate-ascending under upper lip of corolla (except Nepetamultifida. N. tenuifolia, and N. annua where posterior stamens are slightly exserted).   (4)
       
4 (3) Upper lip of corolla 2-plicate inside, plicae parallel to margin, connected at apex; pedicel flattened   24 Lallemantia
+ Upper lip of corolla smooth, very rarely 1-plicate inside; pedicel terete.   (5)
       
5 (4) Calyx upper lip 3-lobed, lower 2-lobed; sinuses of teeth without a thickened fold.   (6)
+ Calyx upper lip 3-toothed and lower lip 2-toothed or upper lip entire and lower lip 4-toothed; sinuses of some or all teeth with a thickened fold   23 Dracocephalum
       
6 (5) Anther cells divaricate at an angle of nearly 180°; plants without stolons; calyx 11-15-veined, with a truncate or oblique mouth; nutlets warty or smooth   18 Nepeta
+ Anther cells divergent or parallel; plants always stoloniferous; calyx 2-lipped, upper lip 3-toothed, lower lip 2-toothed.   (7)
       
7 (6) Anther cells diverging at right angles; leaves mostly obtuse or rounded at apex; flowers less than 3 cm.   (8)
+ Anther cells parallel; leaves acute or short acuminate at apex; plants stoloniferous or not; flowers generally more than 3 cm   22 Meehania
       
8 (7) Calyx tubular or campanulate, 15-veined; teeth narrowly triangular to ovate-triangular or ovate, always awned or awned-spinescent at apex; corolla tube not pilose annulate inside; nutlets less than 1.5 mm   19 Glechoma
+ Calyx obconical-campanulate, 5-veined; teeth ovate-deltoid, acute, or acuminate, slightly curved at apex; corolla pilose annulate near base inside; nutlets ca. 2 mm   20 Suzukia

List of Keys

  • List of lower taxa


     

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