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101. Calanthe R. Brown, Bot. Reg. 7: ad t. 573 ["578"]. 1821.

虾脊兰属 xia ji lan shu

Authors: Xinqi Chen, Phillip J. Cribb & Stephan W. Gale

Alismorkis Thouars.

Herbs, terrestrial, small to large. Rhizome present or not. Pseudobulbs usually small, conic, ovoid, or subglobose, rarely inconspicuous or cylindric, with roots at base and a few leaves at apex; roots long, stout, pubescent. Leaves sometimes not well developed or not completely spreading at anthesis; blade rarely narrowly ensiform or lorate, ± plicate, but often convolute when young, base contracted into a petiole-like stalk or subsessile, sheathing, articulate or not. Scape arising from leaf axil or from base of pseudobulb, rarely from apex of leafless pseudobulb, erect, usually densely puberulent; inflorescence a terminal raceme, few to many flowered; floral bracts persistent or caducous. Flowers resupinate, small to medium-sized, often opening widely. Sepals similar, free. Petals often smaller than sepals; lip adnate to base of column wings and forming a tube, or adnate only to base of column, or to column foot and free from column itself, lobed or unlobed, base spurred or not; disk often lamellate or ridged, sometimes with callose appendages at base. Column often short and thick, without or with a short foot at base, winged, rostellum lobed or unlobed; stigma lateral; pollinia waxy, 8, in 2 groups, nearly equal or unequal in size; caudicle conspicuous or inconspicuous, commonly adherent to a sticky viscidium.

About 150 species: tropical and subtropical Asia, Australia, New Guinea, and the SW Pacific islands, as well as tropical Africa and Central and NW South America; 51 species (21 endemic) in China.


1 Leaves deciduous in dry season; inflorescence (including rachis, bracts, and flowers) densely villous; column with a distinct column foot; lip adnate to column foot.   51 C. labrosa
+ Leaves not deciduous; inflorescence glabrous or puberulent; column without a column foot; lip adnate to column wings or adnate to base of column   (2)
       
2 (1) Floral bracts caducous; rostellum unlobed   (3)
+ Floral bracts persistent; rostellum 2- or 3-lobed   (8)
       
3 (2) Lip similar to petals, spurless.   1 C. actinomorpha
+ Lip different from petals, spurred at base   (4)
       
4 (3) Flowers white.   2 C. angustifolia
+ Flowers yellow   (5)
       
5 (4) Lip obscurely 3-lobed; lateral lobes very small, to 1 mm, sometimes inconspicuous.   3 C. lyroglossa
+ Lip distinctly 3-lobed; lateral lobes relatively large, 2-5 mm   (6)
       
6 (5) Spur cylindric; lip adnate to base of column wings; column slender, ca. 12 mm.   4 C. densiflora
+ Spur clavate; lip adnate to entire length of column wings; column thick, 5-7 mm   (7)
       
7 (6) Leaves narrowly elliptic; lip with 2 triangular lamellae at base of disk.   5 C. clavata
+ Leaves oblong-elliptic to oblanceolate; lip with 2 lunar calli between lateral lobes of disk.   6 C. speciosa
       
8 (2) Lip spurless   (9)
+ Lip spurred   (14)
       
9 (8) Flowers small; sepals to 7 mm.   7 C. tsoongiana
+ Flowers large; sepals 10-20 mm   (10)
       
10 (9) Lip clawed at base, claw forming a shallow sac at base.   8 C. sacculata
+ Lip not clawed   (11)
       
11 (10) Lip with 3-5 crested lamellae on disk.   9 C. tricarinata
+ Lip without lamellae or calli on disk   (12)
       
12 (11) Flowers pale yellow.   10 C. ecarinata
+ Flowers pink or pale purplish pink   (13)
       
13 (12) Sepals and petals not reflexed.   11 C. puberula
+ Sepals and petals strongly reflexed.   12 C. reflexa
       
14 (8) Lip unlobed   (15)
+ Lip 2- or 3-lobed   (19)
       
15 (14) Lip margin fimbriate.   13 C. alpina
+ Lip margin entire, undulate, erose, or emarginate, but not fimbriate   (16)
       
16 (15) Floral bracts and sepals reflexed.   14 C. whiteana
+ Floral bracts and sepals not reflexed   (17)
       
17 (16) Leaves 4 or 5, narrowly oblong, 1.5-2.6 cm wide.   15 C. fargesii
+ Leaves 2 or 3, oblong or oblanceolate, 4-8 cm wide   (18)
       
18 (17) Flowers yellowish green; lip reniform; spur 11-14 mm.   16 C. simplex
+ Flowers white, tinged purple; lip elliptic; spur ca. 28 mm.   17 C. petelotiana
       
19 (14) Lip 2-lobed.   18 C. biloba
+ Lip 3-lobed   (20)
       
20 (19) Lip adorned with wartlike calli on disk   (21)
+ Lip adorned with ridges or lamellae, or without any appendages on disk   (30)
       
21 (20) Mid-lobe of lip entire or shallowly emarginate, mucronate at apex or not   (22)
+ Mid-lobe of lip divided by a deep sinus into 2 lobules   (24)
       
22 (21) Flowers yellow-green; spur ca. 5.5 mm.   19 C. dulongensis
+ Flowers pinkish purple or purple-red; spur 25-50 mm   (23)
       
23 (22) Leaves densely pubescent on both surfaces; pedicel and ovary ca. 20 mm; lip 8-9 mm.   20 C. sinica
+ Leaves sparsely pubescent abaxially; pedicel and ovary ca. 35 mm; lip 11-17 mm.   21 C. sylvatica
       
24 (21) Scape arising from previous year’s leafless pseudobulb.   23 C. odora
+ Scape arising from leaf axil of current year’s growth   (25)
       
25 (24) Spur 2-3 mm.   22 C. ×dominyi
+ Spur 5-30 mm   (26)
       
26 (25) Leaves ensiform or lorate, 0.8-2(-5) cm wide; floral bracts reflexed; dorsal sepal to 9 mm.   24 C. davidii
+ Leaves elliptic, elliptic-lanceolate, ovate-elliptic, or ovate-lanceolate, 4-12 cm wide; floral bracts not reflexed; dorsal sepal 10-15 mm   (27)
       
27 (26) Flowers yellowish green; lateral lobes of lip ovate or flabellate, 5-7 mm wide   (28)
+ Flowers white, sometimes tinged purplish violet, or occasionally purplish red; lateral lobes of lip linear, oblong, or narrowly obovate-cuneate, 2-3 mm wide   (29)
       
28 (27) Leaves uniformly green, without silver-gray bands on adaxial surface; spur longer than 20 mm.   25 C. herbacea
+ Leaves with several silver-gray bands on adaxial surface; spur less than 20 mm.   26 C. argenteostriata
       
29 (27) Pedicel and ovary 10-20 mm; lobules of lip mid-lobe subrhombic, 5-6 mm wide.   27 C. alismatifolia
+ Pedicel and ovary 30-40 mm; lobules of lip mid-lobe linear-oblong, ca. 3 mm wide.   28 C. triplicata
       
30 (20) Lip without any appendages on disk   (31)
+ Lip adorned with ridges or lamellae on disk (sometimes inconspicuous in C. arisanensis and C. plantaginea)   (32)
       
31 (30) Mid-lobe of lip obovate, ca. 8 mm wide.   29 C. yuana
+ Mid-lobe of lip oblong-ligulate, ca. 2 mm wide.   30 C. limprichtii
       
32 (30) Ovary and capsule with 6 longitudinal winglike ridges.   46 C. arisanensis
+ Ovary and capsule without longitudinal ridges   (33)
       
33 (32) Ridges on lip connecting to column wings   (34)
+ Ridges and lamellae on lip free from column wings   (37)
       
34 (33) Lip with 3-5(-7) ridges on disk; spur 20-32 mm.   45 C. aristulifera
+ Lip with 3 ridges on disk; spur 5-20 mm   (35)
       
35 (34) Lip with 4 brown marks on disk; sepals 12-13 mm.   43 C. graciliflora
+ Lip lacking brown marks on disk; sepals 15-17 mm   (36)
       
36 (35) Ridges on disk of lip inconspicuous; lip mid-lobe as broad as or broader than long.   42 C. plantaginea
+ Ridges on disk of lip tall and conspicuous, running onto mid-lobe; lip mid-lobe much longer than broad.   44 C. henryi
       
37 (33) Lateral lobes of lip distinctly broader than mid-lobe   (38)
+ Lateral lobes of lip narrower than mid-lobe   (40)
       
38 (37) Petiole-like leaf base 30-37 cm; lip ca. 5.5 mm.   39 C. trifida
+ Petiole-like leaf base 3-9 cm; lip 10-13 mm   (39)
       
39 (38) Spur much shorter than pedicel and ovary, 5-10 mm; lip mid-lobe emarginate.   40 C. discolor
+ Spur longer than pedicel and ovary, 20 mm or more; lip mid-lobe shortly apiculate.   41 C. alleizettei
       
40 (37) Spur 8-15 mm   (41)
+ Spur to 6 mm   (44)
       
41 (40) Flowers bright yellow; lip with 5 ridges on disk.   50 C. sieboldii
+ Flowers white, pink, or pale red; lip with 3 lamellae or ridges on disk   (42)
       
42 (41) Margins of lip mid-lobe fimbriate.   47 C. metoensis
+ Margins of lip mid-lobe slightly erose or emarginate, but not fimbriate   (43)
       
43 (42) Leaf solitary; flowers pale red; lip mid-lobe broadly ovate-cuneate, ca. 10 × 10 mm.   48 C. lechangensis
+ Leaves 2; flowers white; lip mid-lobe obovate, ca. 7 × 8 mm.   49 C. nankunensis
       
44 (40) Dorsal sepal 7-9 mm.   31 C. mannii
+ Dorsal sepal 12-35 mm   (45)
       
45 (44) Leaves 0.7-3.5 cm wide   (46)
+ Leaves 5-12 cm wide   (48)
       
46 (45) Flowers with purplish red striations; lip with 7 crested lamellae on disk.   36 C. emeishanica
+ Flowers without striations; lip with 3-5 ridges or lamellae   (47)
       
47 (46) Mid-lobe of lip elliptic-rhombic, 8-10 × 6-7 mm, apex aristate.   37 C. arcuata
+ Mid-lobe of lip suboblong, ca. 6 × 4.5 mm, apex acute.   38 C. nipponica
       
48 (45) Lip with a single lamella on mid-lobe.   35 C. griffithii
+ Lip with 3 lamellae on mid-lobe   (49)
       
49 (48) Lateral lobes of lip broader than long; spur adnate to ovary, 4.5-6 mm; lip disk lamellae taller than long.   34 C. fugongensis
+ Lateral lobes of lip longer than broad; spur free from ovary, 2-3 mm; lip disk lamellae longer than tall   (50)
       
50 (49) Sepals and petals buff colored, with pinkish red striations; lip 6-10 mm wide.   32 C. brevicornu
+ Sepals and petals yellow; lip 14-15 mm wide.   33 C. hancockii

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