1 |
Heads discoid or disciform |
|
(2) |
+ |
Heads radiate |
|
(18) |
|
|
|
|
2 (1) |
Heads disciform |
|
(3) |
+ |
Heads discoid |
|
(5) |
|
|
|
|
3 (2) |
Cauline leaf margins entire or spinulose-serrate; heads borne singly or 2–3 in ± corymbiform arrays; cypselae subcylindric-fusiform, 3–4-angled, ± compressed, with 10–12 faint nerves; pappi of 15–60 rigid, unequal, smooth bristles in 1 series |
|
201 Pyrrocoma (in part), p. 413 |
+ |
Cauline leaf margins entire, dentate, or pinnatifid (lobed); heads in corymbiform arrays; cypselae oblong or oblong-obovoid to elliptic or obovoid, ± compressed or flattened, 2(–4)-nerved or ± nerved on edges; pappi of outer setae or scales plus 5–40 bristles, or of 30–40+ bristles in 2 series (outer usually shorter) |
|
(4) |
|
|
|
|
4 (3) |
Plants ± densely white-tomentose (at least some surfaces); phyllaries 1-nerved, nerves not golden-resinous; disc corollas yellowish, throats narrowly funnelform, not indurate, veins light, not resinous; pappi of 30–40+ bristles in 2 series (outer usually shorter) |
|
142 Laënnecia (in part), p. 36 |
+ |
Plants ± hirsute or pilose; phyllaries usually 1–3-nerved, nerves golden-resinous; disc corollas yellow, strongly constricted basally, throats sometimes strongly inflated-indurate, nerves often orange-resinous; pappi of outer setae (or 0) plus 7–25 inner bristles |
|
186 Erigeron (in part), p. 256 |
|
|
|
|
5 (2) |
Disc corollas white or pale yellow; pappi of 10 basally connate scales |
|
153 Thurovia (in part), p. 86 |
+ |
Disc corollas yellow (white in Brintonia); pappi usually of distinct bristles or of outer, shorter setae or scales plus inner, longer bristles, sometimes 0 |
|
(6) |
|
|
|
|
6 (5) |
Stems and leaves resinous, gland-dotted; phyllary apices hooked or looped, or slightly recurved to straight; pappi of 2–8 bristles |
|
202 Grindelia (in part), p. 424 |
+ |
Stems and leaves sometimes resinous, sometimes stipitate-glandular or gland-dotted; phyllary apices straight to squarrose; pappi of 10+ bristles |
|
(7) |
|
|
|
|
7 (6) |
Phyllary midveins translucent and swollen at least basally (not resinous) |
|
(8) |
+ |
Phyllary midveins usually not notably swollen (orange-resinous in Erigeron) |
|
(9) |
|
|
|
|
8 (7) |
Disc corollas white |
|
162 Brintonia, p. 106 |
+ |
Disc corollas yellow |
|
163 Solidago (in part), p. 107 |
|
|
|
|
9 (7) |
Phyllary and corolla nerves orange-resinous; pappi of outer, shorter setae or scales plus inner, longer bristles |
|
186 Erigeron (in part), p. 256 |
+ |
Phyllary and corolla nerves not orange-resinous, or only corolla nerves orange-resinous; pappi of equal or unequal bristles. |
|
(10) |
|
|
|
|
10 (9) |
Phyllaries subequal, foliaceous |
|
169 Toiyabea, p. 172 |
+ |
Phyllaries unequal (sometimes subequal, then outer not foliaceous), outer ± herbaceous (sometimes foliaceous), or chartaceous, or proximally indurate |
|
(11) |
|
|
|
|
11 (10) |
Phyllaries keeled, distally only with relatively small green zones or green along midveins; pappus bristles in 2–3 series (outer in notably shorter series or relatively few) |
|
(12) |
+ |
Phyllaries usually flat, sometimes keeled, distally herbaceous or green; pappi of equal or unequal bristles in 1–3 series (outer not in notably shorter series) |
|
(13) |
|
|
|
|
12 (11) |
Leaves cauline (proximal withering by flowering), margins without coarse spreading cilia near bases; phyllary margins often reddish, sometimes hyaline, abaxial faces glabrous or glabrate to woolly, sometimes stipitate-glandular |
|
143 Eucephalus (in part), p. 39 |
+ |
Leaves basal and cauline (basal and proximal withering by flowering), margins with coarse spreading cilia near bases or on petioles; phyllary margins not reddish, scarious, abaxial faces ± hispid or stipitate-glandular |
|
185 Heterotheca (in part), p. 230 |
|
|
|
|
13 (11) |
Leaf faces densely scabrous and short-stipitate-glandular; phyllaries ± keeled proxi-mally (Esmeralda County, Nevada) |
|
172 Tonestus (in part), p. 181 |
+ |
Leaf faces glabrous or canescent, ± puberulent, hispidulous, tomentose, or villous, and/or sometimes ± stipitate-glandular or gland-dotted; phyllaries sometimes keeled |
|
(14) |
|
|
|
|
14 (13) |
Leaves basal and cauline, basal and proximal petiolate, distal sessile; disc floret style appendages lanceolate |
|
(15) |
+ |
Leaves cauline, sessile; disc floret style appendages triangular |
|
(16) |
|
|
|
|
15 (14) |
Plants densely stipitate-glandular, with caudices; stems densely clustered, simple; leaf blades obovate or oblong to broadly oblanceolate; phyllaries keeled; disc corolla throats cylindric; cypselae white strigoso-hirsute |
|
194 Triniteurybia, p. 382 |
+ |
Plants sparsely, if at all, stipitate-glandular, taprooted; stems single, usually branched; leaf blades lanceolate to oblanceolate; phyllaries flat; disc corolla throats funnelform; cypselae sparsely appressed-hairy |
|
197 Dieteria (in part), p. 395 |
|
|
|
|
16 (14) |
Phyllary margins scarious; disc corolla lobes unequal; cypselae obpyramidal, sericeous; pappi of 40–50 tawny bristles |
|
205 Isocoma (in part), p. 439 |
+ |
Phyllary margins not scarious; disc corolla lobes equal; cypselae prismatic, narrowly turbinate, fusiform, or deltoid, glabrous or sparsely scabrous; pappi of 15–30 reddish brown to brownish bristles |
|
(17) |
|
|
|
|
17 (16) |
Stems glabrous or scabrous, eglandular; leaf bases not subclasping, margins entire; cypselae prismatic or narrowly turbinate; pappus bristles barbellate |
|
200 Oönopsis (in part), p. 410 |
+ |
Stems scabrous to sparsely tomentulose, distally stipitate-glandular; leaf bases subclasping, margins serrate; cypselae fusiform to deltoid; pappus bristles smooth |
|
206 Hazardia (in part), p. 445 |
|
|
|
|
18 (1) |
Rays usually yellow or orange, sometimes cream-colored or white |
|
(19) |
+ |
Rays white, pink, purple, or blue |
|
(38) |
|
|
|
|
19 (18) |
Pappi 0, or crowns, or of scales |
|
(20) |
+ |
Pappi of bristles, or of outer shorter setae, bristles, or scales plus inner, longer bristles, or of setiform awns. |
|
(21) |
|
|
|
|
20 (19) |
Leaves cauline (basal withering by flowering), blades linear to lanceolate or spatulate, stipitate-glandular; heads borne singly or in clusters of 3–6; phyllaries in 2–4 series; cypselae clavate or cylindric, not compresssed, 5–8-nerved, white-hairy (hairs clavate to bulbous); pappi usually of 5–10 scales, sometimes crowns or 0 |
|
155 Gutierrezia (in part), p. 88 |
+ |
Leaves basal (present at flowering) and cauline, blades spatulate, eglandular; heads borne singly (± sessile); phyllaries in 3–7+ series; cypselae obovoid to lanceoloid, flattened, 2-nerved, hairy (hairs glochidiform); pappi of 10–12 (ray) or 20–35+ (disc), lanceolate to subulate or setiform scales |
|
176 Townsendia (in part), p. 193 |
|
|
|
|
21 (19) |
Pappi of outer shorter setae, bristles, or scales plus inner, longer bristles in 2 series |
|
(22) |
+ |
Pappi of setiform scales or bristles in 1–4 series |
|
(26) |
|
|
|
|
22 (21) |
Plants taprooted or with simple caudices (and fibrous-rooted); heads borne singly or in 2s or 3s; phyllaries in 2–3 series, equal or subequal, flat, usually 1–3-nerved (nerves golden-resinous); cypselae 2-nerved, not resinous |
|
186 Erigeron (in part), p. 256 |
+ |
Plants taprooted, rhizomatous, or with branched caudices; heads in corymbiform, ± paniculiform, or subumbelliform arrays, sometimes borne singly; phyllaries in 3–5 series, unequal, usually thickened or keeled (not in Bradburia), 1-nerved, not golden-resinous; cypselae smooth or 1–14-nerved or -ribbed, nerves or ribs sometimes resinous |
|
(23) |
|
|
|
|
23 (22) |
Stems and leaves silky-sericeous, rarely glabrate; leaves sessile, blades 3–11-parallel-nerved (nerves sunken), linear to lanceolate or ovate (often grasslike), margins entire; involucres turbinate |
|
183 Pityopsis, p. 222 |
+ |
Stems and leaves whitish-strigose, pilose, hispid, or arachnose to woolly (hairs flagelliform, soft), or glabrous; leaves sessile or basal petiolate, blades usually 1-nerved (veins reticulate, raised abaxially), spatulate, ovate-oblanceolate, ovate, elliptic, elliptic-oblong, oblanceolate, linear-lanceolate, or (usually distal) linear, margins entire, serrate, or dentate (sometimes coarsely ciliate); involucres campanulate or turbinate |
|
(24) |
|
|
|
|
24 (23) |
Plants proximally woolly, distally arachnose or pilose (hairs flagelliform); basal leaves sessile |
|
182 Chrysopsis (in part), p. 213 |
+ |
Plants sparsely to ± densely hispid, strigose, or pilose (hairs not flagelliform); basal leaves petiolate |
|
(25) |
|
|
|
|
25 (24) |
Perennials (taproots relatively short and/or caudices woody); basal petioles ciliate; cauline leaves much reduced distally, not clasping; heads borne singly or in lax paniculiform arrays; cypselae monomorphic, ray pappi of outer, shorter, setiform scales or bristles plus inner, longer bristles |
|
181 Bradburia (in part), p. 211 |
+ |
Perennials (caudices woody); basal petioles long-strigoso-ciliate; cauline leaves ± reduced distally, sometimes clasping or subclasping; heads borne singly or in corymbiform, sometimes paniculiform arrays; cypselae dimorphic or monomorphic, ray pappi 0, or of outer, linear-lanceolate scales plus inner, longer bristles |
|
185 Heterotheca (in part), p. 230 |
|
|
|
|
26 (21) |
Stems and leaves resinous, gland-dotted; involucres usually globose, hemispheric, or broadly urceolate, sometimes campanulate to obconic, usually ± resinous; pappi of (1–)2–8 white, straight, contorted, or curled, barbellate or smooth awns or setiform scales in 1 series |
|
202 Grindelia (in part), p. 424 |
+ |
Stems and leaves usually not resinous, sometimes stipitate-glandular or gland-dotted; involucres hemispheric, campanulate, turbinate, or cylindric, not resinous, sometimes viscid or stipitate-glandular or gland-dotted; pappi of white, tawny, brownish, or reddish, barbellate bristles in 1–4 series |
|
(27) |
|
|
|
|
27 (26) |
Plants ± colonial (rhizomatous); heads usually glomerate, and/or sometimes pedunculate-solitary in flat-topped or multi-storied, corymbiform arrays |
|
158 Euthamia (in part), p. 97 |
+ |
Plants colonial (sometimes rhizomatous, stems single or clustered, usually from ± woody taproots or caudices); heads borne singly, or (2–5) in paniculiform to corymbiform arrays, or in open or dense corymbiform, paniculiform (diversely shaped in Solidago), racemiform, or spiciform arrays (if glomerate, not in flat-topped, corymbiform arrays, arrays not storied) |
|
(28) |
|
|
|
|
28 (27) |
Plants rhizomatous (often colonial); heads in dense corymbiform or paniculiform arrays |
|
(29) |
+ |
Plants usually taprooted or with caudices, sometimes also from spreading roots (stems single or clustered); heads usually borne singly, sometimes (2–5) in loose corymbiform, cymiform, or paniculiform arrays, or in spiciform, racemiform, paniculiform, or corymbiform arrays, or glomerate (some Oönopsis) |
|
(30) |
|
|
|
|
29 (28) |
Stems and leaves sometimes stipitate-glandular or gland-dotted; cauline leaf bases not clasping, faces sometimes resinous; heads in rounded, club-shaped, wand-shaped, or pyramid-shaped paniculiform (often secund) arrays, or in flat-topped, corymbiform arrays; involucres campanulate to cylindric, 3–12 mm, sometimes stipitate-glandular; phyllary midveins usually swollen, translucent, apices often with green zone, sometimes reflexed; ray florets 3–15(–21); disc corolla lobes lanceolate, erect to reflexed, style-branch appendages triangular; cypselae obconic, terete or ± compressed, 5–8-nerved; pappi of unequal bristles in 1–2(–3) series |
|
163 Solidago (in part), p. 107 |
+ |
Stems and leaves stipitate-glandular; cauline leaf bases clasping or subclasping, faces resinous; heads in dense, flat-topped, corymbiform arrays; involucres campanulate to hemispheric, 10–11 mm, stipitate-glandular; phyllary midveins not swollen, apices green-tipped and spreading-reflexed; ray florets 12–20; disc corolla lobes triangular, spreading, style-branch appendages linear; cypselae fusiform, distinctly compressed, 12–16-nerved (nerves whitish, raised); pappi of equal bristles in 2(–3) series; w United States |
|
164 Oreochrysum, p. 166 |
|
|
|
|
30 (28) |
Stems sometimes clustered; heads usually in rounded, club-shaped, wand-shaped, or pyramid-shaped, paniculiform, or flat-topped, corymbiform (often secund) arrays, or in spiciform, racemiform, or cymiform arrays, sometimes borne singly |
|
(31) |
+ |
Stems single or clustered; heads borne singly or (2–5) in loose, corymbiform or paniculiform arrays, or 2–5(–15) in racemiform, spiciform, or corymbiform arrays |
|
(32) |
|
|
|
|
31 (30) |
Plants with caudices (stems clustered); leaves basal and cauline; heads in rounded, club-shaped, wand-shaped, or pyramid-shaped, paniculiform (often secund) or flat-topped corymbiform arrays; involucres campanulate to cylindric, 3–12 mm; phyllary midveins usually swollen and translucent; cypselae 5–8-nerved |
|
163 Solidago (in part), p. 107 |
+ |
Plants taprooted; leaves mostly cauline; heads usually in spiciform, racemiform, or cymiform arrays, sometimes borne singly; involucres campanulate, 11–13 mm; phyllary midveins not swollen; cypselae 4–5-nerved (California, Oregon) |
|
206 Hazardia (in part), p. 445 |
|
|
|
|
32 (30) |
Stems single, usually branched; cypselae dimorphic (rays 3-angled, discs ± compressed), obovoid, oblong, or obscurely cordate, or rays broadly ellipsoid to obovoid (thick-walled) and discs broadly ellipsoid to clavate (thin-walled); pappi of coarse, barbellate bristles or setiform scales |
|
(33) |
+ |
Stems single or clustered, usually simple; cypselae monomorphic, terete or compressed to ± flattened, prismatic (3–4-angled), narrowly turbinate, subcylindric-fusiform, cylindric, or linear (usually thin-walled); pappi of relatively fine barbellate bristles |
|
(34) |
|
|
|
|
33 (32) |
Basal leaves 1–2-pinnatifid, cauline deeply incised to coarsely dentate or ± entire; involucres 6–10 mm; phyllaries stiff, leathery or bases indurate, apices herbaceous; disc corolla throats not abruptly ampliate, style-branch append-ages lanceolate |
|
195 Xanthisma (in part), p. 383 |
+ |
Leaves linear to oblanceolate, coarsely serrate to entire; involucres 7–16 mm; phyllaries herbaceous; disc corolla throats abruptly ampliate, style-branchappendages deltate |
|
204 Rayjacksonia (in part), p. 437 |
|
|
|
|
34 (32) |
Leaves 3-nerved, faces arachnose (in minute, abaxial lacunae); involucresnarrowly turbinate to subcylindric; pappi tawny to reddish brown |
|
184 Croptilon (in part), p. 228 |
+ |
Leaves 1–3(–5)-nerved, faces glabrous, scabrous, villous, sericeous, tomentose, or lanate; involucres hemispheric to broadly or narrowly campanulate; pappi whitish or brownish |
|
(35) |
|
|
|
|
35 (34) |
Plants sometimes mat-forming; stems clustered; leaves 1–5-nerved; margins of outer phyllaries scarious; pappi whitish |
|
(36) |
+ |
Plants not mat-forming; stems single or clustered; leaves 1-nerved; margins of outer phyllaries herbaceous (sometimes proximally indurate); pappi brownish |
|
(37) |
|
|
|
|
36 (35) |
Stems eglandular or stipitate-glandular, sometimes resinous; leaf margins entire, faces glabrous, scabrous, villous, or lanate, usually stipitate-glandular, sometimes eglandular; phyllaries unequal, proximally indurate, distally chartaceous or herbaceous; ray florets 5–17; cypselae usually sericeous, sometimes glabrous |
|
170 Stenotus, p. 174 |
+ |
Stems densely stipitate-glandular (viscid); leaf margins entire, coarsely and irregularly serrate, dentate, or lobed, faces glabrous or scabrous, sometimes stipitate-glandular; phyllaries subequal, outer foliaceous; ray florets 11–23(–35); cypselae glabrous or villous |
|
172 Tonestus (in part), p. 181 |
|
|
|
|
37 (35) |
Stems and leaves glabrous or sparsely tomentose, eglandular; leaves basal (usually withering by flowering) and cauline, blades narrowly oblanceolate to lanceolate or linear, margins entire; heads borne singly, or 2–12 in glomerules or in loose, corymbiform arrays (subtended by little-reduced distal leaves); rays 6–25; cypselae prismatic or narrowly turbinate |
|
200 Oönopsis (in part), p. 410 |
+ |
Stems and leaves loosely tomentose to woolly, sometimes gland-dotted or stipitate-glandular; leaves basal (persistent) and cauline, basal blades oblanceolate to elliptic or linear, cauline lanceolate, margins entire, spinulose-dentate or -serrate, or shallowly laciniate; heads borne singly or (2–15, ± sessile) in racemiform, spiciform, or loose, corymbiform arrays (at ends of scapiform stems or peduncles); rays 10–80; cypselae subcylindric-fusiform |
|
201 Pyrrocoma (in part), p. 413 |
|
|
|
|
38 (18) |
Receptacles conic; rays white (pink or purplish-tinged abaxially) |
|
(39) |
+ |
Receptacles flat or convex; rays white, pink, blue, or purple (not tinged abaxially) |
|
(40) |
|
|
|
|
39 (38) |
Plants 5–20 cm (scapiform, short-rhizomatous); leaves mostly basal, margins crenate-serrate; phyllary margins herbaceous; rays in 3–4 series (closing at night); cypselae obconic, compressed, margins thickened, riblike, apices not cupped; pappi 0 |
|
139 Bellis, p. 22 |
+ |
Plants 5–50 cm (leafy, taprooted); leaves basal and cauline, margins entire or toothed to pinnatifid; phyllary margins broadly scarious; rays in 1(–2) series (not closing at night); cypselae columnar, usually 4-angled, 4–12-ribbed, apices usually cupped; pappi of awns, aristate scales, short bristles, or 0 |
|
188 Aphanostephus (in part), p. 351 |
|
|
|
|
40 (38) |
Plants colonial; stems branched (lateral branches strongly ascending, commonly modified to green thorns); leaves early withering; phyllaries (1–)3(–5)-nerved (usually wet sites in arid, sw United States) |
|
190 Chloracantha (in part), p. 358 |
+ |
Plants sometimes colonial; stems single or clustered, simple or branched (not becoming thorny); at least distal leaves persistent through flowering; phyllaries usually 1-nerved, seldom 3-nerved |
|
(41) |
|
|
|
|
41 (40) |
Cypselae usually obconic or obovoid, sometimes lanceoloid, flattened or compressed, margins ribbed (sometimes 1–2 nerves also on faces) |
|
(42) |
+ |
Cypseale ± narrowly obconic, obovoid, oblanceoloid, lanceoloid, fusiform, or linear, ± compressed or terete, sometimes winged or wings reduced and resembling marginal nerves or ribs (then pappi 0 or of minute scales), usually 3–12+-nerved on faces (margins not ribbed) |
|
(46) |
|
|
|
|
42 (41) |
Leaves cauline; phyllaries keeled |
|
(43) |
+ |
Leaves basal and/or cauline; phyllaries not keeled |
|
(44) |
|
|
|
|
43 (42) |
Plants 10–160 cm, minutely stipitate-glandular distally; proximalmost leaves scalelike, proximal withering by flowering, cauline distally increasing in size to mid stems, mid and distal blades lanceolate or lance-ovate to elliptic; heads in racemiform or corymbiform arrays; cypselae glabrous, pilose, or strigose, eglandular; pappi of outer, shorter plus inner, longer bristles in 3 series |
|
143 Eucephalus (in part), p. 39 |
+ |
Plants 4–30(–70) cm, sometimes stipitate-glandular; leaves persistent to flowering, mostly equal in size and shape, blades spatulate (proximal), linear, narrowly oblong, or elliptic-lanceolate; heads borne singly or in 2s or 3s, or in corymbiform arrays; cypselae densely strigose or sericeous, sometimes stipitate-glandular; pappi of outer shorter bristles or scales plus inner, longer bristles in 2 series |
|
150 Ionactis (in part), p. 82 |
|
|
|
|
44 (42) |
Leaf faces eglandular; corolla lobes lanceolate; pappi of 20–30 bristles |
|
138 Aster, p. 20 |
+ |
Leaf faces often gland-dotted; corolla lobes deltate or lance-deltate; pappi usually of 12–35+ scales, or of outer setae or scales plus 5–40(–50) bristles (sometimes only on rays or on discs), sometimes 0 |
|
(45) |
|
|
|
|
45 (44) |
Plants taprooted or with branched caudices; heads borne singly; leaves usually entire, rarely toothed or lobed; phyllaries unequal, 1-nerved (nerves not golden-resinous); disc corolla throats funnelform; cypselae glabrous or hairy (hairs glochidiform); pappi of 12–35+ scales |
|
176 Townsendia (in part), p. 193 |
+ |
Plants rhizomatous, sometimes taprooted; heads borne singly or in corymbiform arrays; leaves entire, ± dentate, or pin-natifid; phyllaries equal to unequal, 1–3-nerved (nerves golden-resinous); disc corolla throats sometimes strongly inflated-indurate; cypselae glabrous, strigose, or sericeous; pappi usually of outer, shorter setae or scales plus 5–40(–50) inner, longer bristles (sometimes only on rays or discs),sometimes 0 |
|
186 Erigeron (in part), p. 256 |
|
|
|
|
46 (41) |
Pappi 0, or crowns of setae or scales, or of scales, of alternating scales and bristles, or of outer, shorter setae or scales plus inner, longer bristles |
|
(47) |
+ |
Pappi of bristles or setiform scales |
|
(50) |
|
|
|
|
47 (46) |
Phyllary midnerves not swollen, not orange-resinous; disc style-branch appendages triangular or lanceolate |
|
(48) |
+ |
Phyllary midnerves (or 1–3 nerves) orange-resinous; disc style-branch appendages deltate |
|
(49) |
|
|
|
|
48 (47) |
Leaves usually entire, sometimes dentate or lobed, bases clasping or subclasping; phyllaries subequal; pappi rings or crowns of setae or scales |
|
177 Astranthium (in part), p. 203 |
+ |
Leaves entire, bases not clasping; phyllaries unequal; pappi thickened rings, crowns of scales (basally fused or distinct), or minute outer scales plus inner bristles |
|
179 Chaetopappa (in part), p. 206 |
|
|
|
|
49 (47) |
Plants not colonial (stems ± clustered), stems and leaves stipitate-glandular; basal leaves persistent or withering by flowering; heads borne singly or (2–10) in corymbiform arrays; pappi usually of outer setae or scales plus 5–40(–50) inner bristles, sometimes 0 |
|
186 Erigeron (in part), p. 256 |
+ |
Plants colonial, stems and leaves eglandular; basal leaves withering by flowering; heads in corymbiform or diffuse, paniculiform arrays; pappi of 2–3 awns plus shorter bristles or scales, or wholly of minute scales |
|
189 Boltonia, p. 353 |
|
|
|
|
50 (46) |
Cypselae ± dimorphic (ray 3-sided and rounded abaxially, disc ± compressed); pappi of relatively coarse (± flattened) bristles or setiform scales |
|
(51) |
+ |
Cypselae monomorphic; pappi of relatively fine bristles |
|
(52) |
|
|
|
|
51 (50) |
Stems simple; leaves mostly basal, margins serrate or serrulate; involucres depressed-hemispheric; cypselae 3–9-ribbed on each face; pappus bristles coarsely barbellate |
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195 Xanthisma (in part), p. 383 |
+ |
Stems usually branched ; leaves basal (persistent in rosettes or withering) and cauline (distally ± reduced) or mostly cauline, margins pinnately lobed or pinnatifid, toothed, or entire; involucres turbinate, campanulate, or hemispheric; cypselae 8–13-nerved on each face; pappus bristles barbellulate |
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198 Arida (in part), p. 401 |
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52 (50) |
Plants mostly rhizomatous, sometimes with caudices; heads in paniculiform or racemiform arrays, rarely borne singly |
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(53) |
+ |
Plants taprooted, with caudices, or rhizomatous; heads in corymbiform (or flat-topped racemiform) arrays or borne singly, or sometimes grouped into loose corymbiform arrays |
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(56) |
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53 (52) |
Stems spreading-hirsute, eglandular; heads in narrow (wand-shaped), paniculiform arrays; phyllary midribs translucent and swollen; rays 7–9, corollas white to pale cream |
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163 Solidago (in part), p. 107 |
+ |
Stems usually glabrous, often distally hairy in lines, sometimes ± densely hairy, sometimes distally stipitate-glandular; heads in ± open or dense (not wand-shaped), paniculiform arrays; phyllary midnerves not translucent and swollen; rays (8–)12–65(–75), corollas white, pink, blue, or purple |
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(54) |
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54 (53) |
Phyllaries usually unequal, sometimes subequal, proximally indurate, distally with defined green zone, sometimes distally foliaceous, or outer whollyfoliaceous, sometimes short-stipitate-glandular |
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214 Symphyotrichum (in part), p. 465 |
+ |
Phyllaries subequal, herbaceous (without definite distal green zones, not foliaceous), short-stipitate-glandular |
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(55) |
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55 (54) |
Stems ± densely villous; leaves cauline, blades 1-nerved (venation reticulate), lanceolate to elliptic, bases auriculate-clasping, abaxial faces glabrate to ± strigose, adaxial sparsely villous (distal stipitate-glandular); phyllaries often purplish, apices of outer acuminate; disc corollas whitish to pale yellow; cold wet soils, montane (n Cordilleras) and boreal North America |
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210 Canadanthus, p. 458 |
+ |
Stems glabrous; leaves basal and cauline, blades 3-nerved (nerves ± parallel), linear, bases subclasping, faces glabrous (distal stipitate-glandular); phyllaries green, apices of outer acute; disc corollas yellow; damp alkaline areas, deserts and dry prairies, w North America |
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212 Almutaster, p. 461 |
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56 (52) |
Plants usually tapooted, sometimes with caudices (also rhizomatous from fibrous roots in Chaetopappa); stems usually 1 (sometimes 2–5+ in clusters), branched or simple; heads borne singly or in loose corymbiform arrays |
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(57) |
+ |
Plants rhizomatous or with caudices; stems 1–5+, usually simple; heads usually in corymbiform arrays, sometimes borne singly (then plants long-rhizomatous, rays pink) |
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(62) |
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57 (56) |
Stems and leaves usually densely white-tomentose, sometimes glabrate, stipitate-glandular or gland-dotted; stems simple; ray florets neuter; cypselae cuneiform or linear; pappi reddish to brownish (bristles relatively coarse, California) |
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208 Corethrogyne (in part), p. 450 |
+ |
Stems and leaves glabrous, glabrate, canescent, villous, or tomentose, sometimes stipitate-glandular or gland-dotted; stems simple or branched; ray florets pistillate, fertile; cypselae fusiform, cylindric, obovoid, or linear; pappi hyaline or white to tawny |
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(58) |
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58 (57) |
Stems simple; leaf margins entire; phyllaries mostly foliaceous (margins sometimes proximally indurate); rays white (maturing or drying bluish or purplish) |
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(59) |
+ |
Stems branched or simple; leaf margins entire or toothed (teeth apiculate, ± spinulose, or spine-tipped); phyllaries usually not foliaceous (proximally white-indurate, distally green or herbaceous); rays white, blue, violet, or purple |
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(60) |
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59 (58) |
Leaves basal and cauline (crowded), blades linear-oblanceolate to lanceolate, 1-nerved; phyllaries not keeled; cypselae 5-nerved |
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179 Chaetopappa (in part), p. 206 |
+ |
Leaves mostly basal (rosettes), blades linear to oblanceolate, 3-nerved; phyllaries often ± keeled; cypselae 5–10-nerved (nerves raised) |
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191 Oreostemma, p. 359 |
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60 (58) |
Stems mostly simple (scapiform); leaves mostly basal (rosettes, often marcescent), margins entire or irregularly serrate (teeth apiculate or ± spinulose); phyllaries squarrose; cypselae 8–10-ribbed (canyons, rock faces, Utah) |
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192 Herrickia (in part), p. 361 |
+ |
Stems mostly branched; leaves basal and cauline, margins entire or toothed (teeth spinose-tipped); phyllaries appressed, spreading, or reflexed; cypselae 4-nerved, 4–6-ribbed, or smooth |
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(61) |
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61 (60) |
Stems mostly single; cauline leaf blades lanceolate to oblanceolate; phyllaries not keeled |
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197 Dieteria (in part), p. 395 |
+ |
Stems clustered; cauline leaf blades spatulate to obovate oroblong, rarely elliptic; phyllaries keeled |
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199 Xylorhiza (in part), p. 406 |
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62 (56) |
Phyllaries flat, not keeled, midveins orange-resinous or swollen and translucent |
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(63) |
+ |
Phyllaries ± rounded, sometimes ± keeled, midveins not swollen |
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(64) |
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63 (62) |
Plants 40–200 cm; leaves basal (not persistent, reduced) and cauline, blades 1-nerved (venation brochidodromous), lanceolate to elliptic; phyllary midveins orange-resinous; rays 2–10(–16) |
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144 Doellingeria, p. 43 |
+ |
Plants 10–40 cm; leaves basal (persistent, well developed) and cauline (reduced), blades usually 1-nerved, sometimes ± 3-nerved (venation reticulate), linear to linear-lanceolate, phyllary midveins swollen, translucent; ray florets 10–20 |
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163 Solidago (in part), p. 107 |
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64 (62) |
Plants rhizomatous and/or with woody caudices (rhizomes with swollen apical buds); heads in ± loose, corymbiform arrays or borne singly (nodding in bud); phyllaries lance-ovate to linear (membranous), proximally not indurate, green along midnerves; cypselae 5–8-nerved (lateral 2 thicker), glabrous, ± densely gland-dotted; e North America |
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149 Oclemena, p. 78 |
+ |
Plants rhizomatous (rhizomes not apically swollen) and/or with caudices; heads in corymbiform arrays (erect in bud); phyllaries ovate, oblong, lanceolate, or linear-lanceolate, proximally indurate, distally with sharply delimited green apical zones; cypselae 7–12(–18)-nerved, glabrous, strigillose, or strigose, eglandular; e, w North America (excluding United States prairies) |
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(65) |
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65 (64) |
Involucres cylindric; ray florets 1–6, white; disc corollas white or cream-colored; cypselae ± densely strigose |
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160 Sericocarpus, p. 101 |
+ |
Involucres cylindro-campanulate or campanulate; rays 5–60, white to purple; disc corollas yellow; cypselae glabrous or ± densely strigillose |
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(66) |
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66 (65) |
Stems and leaves usually stipitate-glandular, sometimes eglandular and glaucous; leaves mostly cauline, entire or spinulose-serrate, glabrous or scabrellous; phyllaries sometimes rounded, usually keeled, apices acuteto long-acuminate; rays 8–27; disc corolla tubes shorter than limbs (w Cordilleras) |
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192 Herrickia (in part), p. 361 |
+ |
Stems and leaves usually eglandular, sometimes stipitate-glandular (e North America only), not glaucous; leaves basal and/or cauline, serrate (teeth sometimes spinulose or spinose, blades then linear, grasslike, se North America) or entire, hairy or glabrous; phyllaries usually rounded, sometimes keeled, apices obtuse to acute; rays 5–60; disc corolla tubes shorter or longer than limbs |
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193 Eurybia, p. 365 |
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