1 |
Leaf blade papery or membranous and deciduous |
|
(2) |
+ |
Leaf blade leathery or thinly leathery and evergreen |
|
(15) |
|
|
|
|
2 (1) |
Petiole 2-8 cm; leaf blade orbicular or orbicular-elliptic to broadly ovate |
|
(3) |
+ |
Petiole less than 2 cm; leaf blade not as above |
|
(4) |
|
|
|
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3 (2) |
Leaf blade more than 9.5 cm, auriculate at base; fruit ovoid, 1.3-1.7 × 1.1-1.3 cm, perianth tube cup-shaped. |
|
1 L. auriculata |
+ |
Leaf blade less than 8 cm, base orbicular or cuneate; fruit globose, 0.5-0.6 cm in diam., perianth tube shallowly discoid. |
|
2 L. populifolia |
|
|
|
|
4 (2) |
Branchlets glabrous |
|
(5) |
+ |
Branchlets covered with hairs |
|
(10) |
|
|
|
|
5 (4) |
Leaf blade glabrous abaxially |
|
(6) |
+ |
Leaf blade hairy abaxially or at least in axils of veins |
|
(8) |
|
|
|
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6 (5) |
Leaf blade elliptic or lanceolate-elliptic; flowers 10-12 per umbel; filaments glabrous; pedicels hairy. |
|
5 L. rubescens |
+ |
Leaf blade lanceolate or oblong; flowers 4-6 per umbel; filaments glabrous or with hairs below middle; pedicels glabrous |
|
(7) |
|
|
|
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7 (6) |
Terminal bud ± hairy; leaf blade ± hairy abaxially when young or in bud. |
|
3 L. cubeba |
+ |
Terminal bud glabrous; leaf blade glabrous on both surfaces even when young or in bud. |
|
4 L. kingii |
|
|
|
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8 (5) |
Young leaves gray-white tomentose abaxially; leaf blade obovate or obovate-elliptic, often larger, 7-11 × 3-5 cm. |
|
8 L. tsinlingensis |
+ |
Young leaves hairy only in axils of veins or on each side of midrib; leaf blade often smaller, 2-5 × 1-3 cm |
|
(9) |
|
|
|
|
9 (8) |
Leaf blade usually obovate or rotund, sometimes glandular-domed in axils of veins; lateral veins 4-6 pairs. |
|
6 L. ichangensis |
+ |
Leaf blade elliptic, elliptic-lanceolate, or elliptic-obovate, not glandular-domed in axils of veins; lateral veins 5-8 pairs. |
|
7 L. chunii |
|
|
|
|
10 (4) |
Branchlets and leaf blade abaxially covered with pubescence or tomentum, indumentum of young branchlets scarcely deciduous, persistent in 2-year-old branchlets |
|
(11) |
+ |
Branchlets and leaf blade abaxially sericeous, indumentum of young branchlets soon deciduous, mostly glabrous in 2-year-old flowering and fruiting branchlets |
|
(13) |
|
|
|
|
11 (10) |
Leaf blade sericeous-pubescent or with white pubescence abaxially; flowers 4-6 per umbel. |
|
11 L. mollis |
+ |
Leaf blade densely gray-yellow tomentose abaxially; flowers 8-14 per umbel |
|
(12) |
|
|
|
|
12 (11) |
Leaf blade ovate, rhombic-ovate, or oblong, 4-15 × 1.5-7 cm; umbel solitary. |
|
9 L. moupinensis |
+ |
Leaf blade elliptic or oblong, 1-13.5 × 3.2-4.5 cm; umbels in pairs seated on common peduncle. |
|
10 L. taronensis |
|
|
|
|
13 (10) |
Young branchlets and leaf blade abaxially shortly gray sericeous; leaf blade lanceolate or obovate-lanceolate. |
|
12 L. pungens |
+ |
Young branchlets and leaf blade long yellow or brown sericeous; leaf blade oblong-lanceolate, obovate-oblong, or obovate |
|
(14) |
|
|
|
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14 (13) |
Leaf blade oblong-lanceolate, apex acuminate; peduncles glabrous. |
|
13 L. sericea |
+ |
Leaf blade obovate or obovate-oblong, acute or obtuse at apex; peduncles hairy. |
|
14 L. veitchiana |
|
|
|
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15 (1) |
Perianth segments imperfect, lacking, or 3 or 4 |
|
(16) |
+ |
Perianth segments 6-8 |
|
(17) |
|
|
|
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16 (15) |
Perianth segments imperfect or lacking; fertile stamens usually 15 or more; fruit globose; leaf blade gray-yellow tomentose or subglabrous abaxially. |
|
15 L. glutinosa |
+ |
Perianth segments 3 or 4; fertile stamens 9; fruit ellipsoid; leaf blade along veins shortly hirsute abaxially. |
|
16 L. morrisonensis |
|
|
|
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17 (15) |
Perianth tube not or slightly enlarged at fruit; fruiting tube complanate or shallowly discoid |
|
(18) |
+ |
Perianth tube very enlarged at fruit; fruit ± surrounded by discoid or cup-shaped fruiting tube |
|
(35) |
|
|
|
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18 (17) |
Leaves verticillate, usually 3-6 per whorl |
|
(19) |
+ |
Leaves opposite or alternate |
|
(20) |
|
|
|
|
19 (18) |
Young branchlets and petioles densely ferruginous tomentose; leaf blade leathery, obovate- lanceolate, 3.5-6.8 × 0.9-2.8 cm; umbels lateral in leaf axils. |
|
17 L. verticillifolia |
+ |
Young branchlets and petioles densely yellow or ferruginous hirsute; leaf blade subleathery, lanceolate or oblanceolate-oblong, 7-25 × 2-6 cm; umbels usually clustered at apex of branchlet. |
|
18 L. verticillata |
|
|
|
|
20 (18) |
Leaves opposite or subopposite (rarely alternate on same tree) |
|
(21) |
+ |
Leaves alternate |
|
(22) |
|
|
|
|
21 (20) |
Leaf blade glabrous or subglabrous abaxially; leaves and buds usually tinged with red after autumn. |
|
19 L. variabilis |
+ |
Leaf blade yellow-brown or ferruginous tomentose abaxially; leaves and buds not tinged with red after autumn. |
|
20 L. lancifolia |
|
|
|
|
22 (20) |
Peduncles and pedicels absent. |
|
21 L. rotundifolia |
+ |
Peduncles and pedicels present, if peduncles absent, pedicels still present |
|
(23) |
|
|
|
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23 (22) |
Perianth segments deciduous at fruit; fruit long ellipsoid or long ovoid to globose |
|
(24) |
+ |
Perianth segments persistent at fruit; fruit globose or subglobose |
|
(29) |
|
|
|
|
24 (23) |
Flowers 2 or 3 per umbel |
|
(25) |
+ |
Flowers 4-6 or more per umbel |
|
(26) |
|
|
|
|
25 (24) |
Leaf blade smaller, 4-11 × 1.5-3.5 cm, yellow-brown pubescent abaxially. |
|
29 L. balansae |
+ |
Leaf blade larger, 9-16.5 × 3-6.5 cm, glabrous abaxially. |
|
30 L. biflora |
|
|
|
|
26 (24) |
Leaf blade long caudate at apex, foveolate on both surfaces; fruit globose. |
|
31 L. beilschmiediifolia |
+ |
Leaf blade acuminate, acute, obtuse, or rounded at apex, not foveolate on both surfaces; fruit long ellipsoid or long ovoid |
|
(27) |
|
|
|
|
27 (26) |
Branchlets and petioles glabrous; leaf blade yellow-brown puberulent abaxially. |
|
32 L. salicifolia |
+ |
Branchlets and petioles hairy; leaf blade ferruginous pubescent, pubescent along veins, or glabrous abaxially |
|
(28) |
|
|
|
|
28 (27) |
Branchlets and petioles ferruginous pubescent; leaf blade abaxially ferruginous pubescent, apex obtuse or rounded. |
|
33 L. monopetala |
+ |
Branchlets and petioles gray appressed pubescent; leaf blade glaucous, glabrous, or pubescent along veins abaxially, apex acuminate. |
|
34 L. albescens |
|
|
|
|
29 (23) |
Peduncles absent; fruiting pedicels stout; persistent perianth segments 6, regular, usually erect |
|
(30) |
+ |
Peduncles present; fruiting pedicels usually slender; persistent perianth segments 2-4, irregular, reflexed |
|
(31) |
|
|
|
|
30 (29) |
Young branchlets and young leaf blade abaxially glabrous or densely gray-yellow villous; leaf blade obovate-elliptic, obovate-lanceolate, oblong, or lanceolate, 4.5-9.5 × 1.4-4 cm. |
|
22 L. coreana |
+ |
Young branchlets gray pubescent and young leaf blade abaxially glaucous and gray- white villous along midrib; leaf blade narrowly lanceolate or lanceolate to elliptic, 10-13 × 2-3.5 cm. |
|
23 L. hupehana |
|
|
|
|
31 (29) |
Branchlets, petioles, and filaments glabrous; fruit larger, 2-3 cm in diam |
|
(32) |
+ |
Branchlets, petioles, and filaments hairy; fruit smaller, less than 1 cm in diam |
|
(33) |
|
|
|
|
32 (31) |
Leaf blade not foveolate on both surfaces and adaxially glabrous but ± hairy abaxially along veins; fruit globose, fruiting pedicel somewhat thickened at apex, as peduncle ± hairy and not verrucose. |
|
24 L. honghoensis |
+ |
Leaf blade foveolate and glabrous on both surfaces; fruit ovoid, fruiting pedicel shallowly cup-shaped at apex, as peduncle glabrous and verrucose. |
|
25 L. cangyuanensis |
|
|
|
|
33 (31) |
Leaf blade abaxially, petioles, and peduncles densely ferruginous tomentose. |
|
28 L. umbellata |
+ |
Leaf blade abaxially, petioles, and peduncles covered with brown pubescence or long gray-yellow pubescence |
|
(34) |
|
|
|
|
34 (33) |
Leaf blade greenish abaxially; umbels 2 in leaf axils at tips of branchlets; fruiting pedicels shorter, less than 2 mm. |
|
26 L. foveola |
+ |
Leaf blade blue-green abaxially; several umbels axillary on shortened branchlets; fruiting pedicels longer, more than 6 mm. |
|
27 L. coelestis |
|
|
|
|
35 (17) |
Some umbels inserted on long or slightly long peduncles in panicles, racemes, or near corymbs |
|
(36) |
+ |
Umbels solitary or fascicled |
|
(43) |
|
|
|
|
36 (35) |
Young branchlets and leaf blade abaxially glabrous |
|
(37) |
+ |
Young branchlets and leaf blade abaxially hairy (only in L. panamanja glabrous abaxially) |
|
(39) |
|
|
|
|
37 (36) |
Leaf blade larger, 21-50 × 11-14.5 cm, lateral veins 15-22 pairs. |
|
35 L. dilleniifolia |
+ |
Leaf blade smaller, 10-21 × 3-8 cm, lateral veins 7-12 pairs |
|
(38) |
|
|
|
|
38 (37) |
Leaf blade midrib prominent adaxially, lateral veins 10-12 pairs; fertile stamens 26-32. |
|
36 L. liyuyingii |
+ |
Leaf blade midrib impressed adaxially, lateral veins 7-9 pairs; fertile stamens 9. |
|
37 L. szemaois |
|
|
|
|
39 (36) |
Leaf blade abaxially and petiole glabrous; fruit compressed globose. |
|
38 L. panamanja |
+ |
Leaf blade abaxially and petiole yellow-brown or ferruginous pubescent (at least young leaves so); fruit oblong or compressed globose |
|
(40) |
|
|
|
|
40 (39) |
Petiole more than 2 cm; perianth segments 8; fertile stamens 12-14; fruit compressed globose. |
|
39 L. semecarpifolia |
+ |
Petiole less than 2 cm; perianth segments 6; fertile stamens 9; fruit oblong |
|
(41) |
|
|
|
|
41 (40) |
Leaf blade obovate to obovate-oblong, rounded, mucronate at apex. |
|
42 L. longistaminata |
+ |
Leaf blade elliptic or elliptic-lanceolate, acuminate, long acuminate, or falcate-acuminate at apex |
|
(42) |
|
|
|
|
42 (41) |
Leaf blade lateral veins 6-8 pairs; fruiting tube deeply cup-shaped. |
|
40 L. martabanica |
+ |
Leaf blade lateral veins 8-12 pairs; fruiting tube shallowly discoid. |
|
41 L. tibetana |
|
|
|
|
43 (35) |
Leaves clustered toward apex of branchlet, subverticillate. |
|
74 L. machiloides |
+ |
Leaves alternate |
|
(44) |
|
|
|
|
44 (43) |
Young branchlets glabrous or subglabrous; petiole usually glabrous when young |
|
(45) |
+ |
Young branchlets and petioles hairy (petioles glabrous in L. baviensis and L. chinpingensis) |
|
(54) |
|
|
|
|
45 (44) |
Leaf blade smaller, mostly less than 9 cm |
|
(46) |
+ |
Leaf blade larger, mostly more than 10 cm |
|
(49) |
|
|
|
|
46 (45) |
Young branchlets glabrous or densely pubescent; leaf blade midrib prominent adaxially; peduncles slender. |
|
43 L. pedunculata |
+ |
Young branchlets glabrous; leaf blade midrib plane or impressed adaxially; peduncles stouter and shorter |
|
(47) |
|
|
|
|
47 (46) |
Leaf blade oblong or narrowly oblong, usually 4 × as long as wide, slightly acute or obtuse at apex. |
|
46 L. kwangtungensis |
+ |
Leaf blade elliptic, obovate, obovate-lanceolate, or lanceolate, ca. 2-3 × as long as wide, rounded-obtuse or shortly acute or acuminate at apex |
|
(48) |
|
|
|
|
48 (47) |
Leaf blade elliptic or rarely obovate, 2 × as long as wide, rounded-obtuse at both ends or base cuneate; fruiting tube cup-shaped. |
|
44 L. pittosporifolia |
+ |
Leaf blade obovate-lanceolate or lanceolate, 3 × as long as wide, shortly acute or acuminate at apex, cuneate or acute at base; fruiting tube discoid. |
|
45 L. hunanensis |
|
|
|
|
49 (45) |
Branchlets with corky cortices or spots |
|
(50) |
+ |
Branchlets without corky cortices or spots |
|
(51) |
|
|
|
|
50 (49) |
Branchlets with corky cortices; leaf blade oblanceolate or narrowly long elliptic; fertile stamens 9; filaments villous. |
|
47 L. suberosa |
+ |
Branchlets with corky spots; leaf blade elliptic; fertile stamens 12(-14); filaments pubescent. |
|
48 L. chengshuzhii |
|
|
|
|
51 (49) |
Fruit compressed globose, 2-2.2 × ca. 3 cm |
|
(52) |
+ |
Fruit ellipsoid or oblong, 1-2.5 × 0.3-1.4 cm |
|
(53) |
|
|
|
|
52 (51) |
Leaf blade lateral veins 6-10 pairs. |
|
49 L. depressa |
+ |
Leaf blade lateral veins 16-20 pairs. |
|
50 L. garciae |
|
|
|
|
53 (51) |
Leaf blade midrib conspicuously prominent on both surfaces; fruit oblong, larger, 1.5-2.5 × 1-1.4 cm; fruiting tube discoid. |
|
51 L. lancilimba |
+ |
Leaf blade midrib impressed adaxially; fruit ellipsoid, smaller, ca. 1.5 × 0.8 cm; fruiting tube cup-shaped. |
|
52 L. subcoriacea |
|
|
|
|
54 (44) |
Young branchlets and petioles puberulent or pubescent, soon deciduous, 2-year-old flowering and fruiting branchlets mostly glabrous |
|
(55) |
+ |
Young branchlets and petioles covered with tomentum or pubescence, indumentum late deciduous, 2-year-old branchlets still covered with more hairs |
|
(63) |
|
|
|
|
55 (54) |
Terminal bud scaly; fruit smaller, less than 1.5 × 1 cm |
|
(56) |
+ |
Terminal bud naked; fruit larger, more than 2 × 1.3 cm |
|
(59) |
|
|
|
|
56 (55) |
Leaf blade glabrous abaxially. |
|
53 L. greenmaniana |
+ |
Leaf blade gray-yellow pubescent or along veins pilose abaxially |
|
(57) |
|
|
|
|
57 (56) |
Young branchlets pubescent; leaf blade pubescent abaxially; lateral veins 5-7 pairs. |
|
54 L. hypophaea |
+ |
Young branchlets puberulent or subglabrous; leaf blade puberulent abaxially; lateral veins 8-10 pairs |
|
(58) |
|
|
|
|
58 (57) |
Leaf blade lanceolate, oblanceolate, or oblong, usually narrower; rudimentary pistil present. |
|
55 L. gongshanensis |
+ |
Leaf blade elliptic, usually broader; rudimentary pistil absent. |
|
56 L. viridis |
|
|
|
|
59 (55) |
Fruiting peduncles and pedicels 2-3 mm |
|
(60) |
+ |
Fruiting peduncles more than 5 mm; fruiting pedicels more than 10 mm |
|
(61) |
|
|
|
|
60 (59) |
Leaf blade lateral veins 7 or 8 pairs; fruiting tube thickly woody, larger, warty outside. |
|
60 L. baviensis |
+ |
Leaf blade lateral veins 10-12 pairs; fruiting tube leathery, smaller, not warty outside. |
|
61 L. litseifolia |
|
|
|
|
61 (59) |
Leaf blade gray puberulent or along veins pubescent abaxially; fruiting tube cup-shaped. |
|
57 L. yunnanensis |
+ |
Leaf blade glabrous abaxially; fruiting tube discoid or cup-shaped |
|
(62) |
|
|
|
|
62 (61) |
Leaf blade lanceolate or narrowly elliptic, usually narrower, less than 4 cm in diam.; lateral veins 7-11 pairs; fruiting tube discoid, not dehiscent. |
|
58 L. chinpingensis |
+ |
Leaf blade elliptic, usually broader, more than 4 cm in diam.; lateral veins 5-7 pairs; fruiting tube cup-shaped, with many lenticels. |
|
59 L. vang |
|
|
|
|
63 (54) |
Leaf blade broadly linear; fruit long ovoid. |
|
62 L. pseudoelongata |
+ |
Leaf blade not broadly linear; fruit not long ovoid |
|
(64) |
|
|
|
|
64 (63) |
Petiole less than 8 mm |
|
(65) |
+ |
Petiole mostly more than 10 mm |
|
(66) |
|
|
|
|
65 (64) |
Leaf blade lateral veins 4 or 5 pairs, impressed abaxially; perianth segments persistent at fruit. |
|
63 L. oligophlebia |
+ |
Leaf blade lateral veins 8-12 pairs, slightly prominent or prominent adaxially; perianth segments deciduous at fruit. |
|
64 L. yaoshanensis |
|
|
|
|
66 (64) |
Leaf blade glabrous or only along veins hairy abaxially |
|
(67) |
+ |
Leaf blade hairy abaxially |
|
(68) |
|
|
|
|
67 (66) |
Leaf blade obovate or oblong-obovate, along veins pubescent abaxially, lateral veins 5-8 pairs. |
|
65 L. akoensis |
+ |
Leaf blade lanceolate or oblong-oblanceolate, glabrous abaxially, lateral veins 8-12 pairs. |
|
66 L. kwangsiensis |
|
|
|
|
68 (66) |
Flowers ca. 20 per umbel. |
|
67 L. liboshengii |
+ |
Flowers 3-6 per umbel |
|
(69) |
|
|
|
|
69 (68) |
Fruit globose or subglobose; leaf buds axillary. |
|
68 L. sinoglobosa |
+ |
Fruit ellipsoid; leaf buds mostly inserted at apex of branchlet |
|
(70) |
|
|
|
|
70 (69) |
Leaf blade obovate or broadly oblong, broader, mostly more than 5 cm in diam |
|
(71) |
+ |
Leaf blade lanceolate, narrowly lanceolate, oblong-lanceolate to oblanceolate, or oblong to lanceolate-elliptic, usually narrower, mostly less than 4 cm in diam |
|
(72) |
|
|
|
|
71 (70) |
Young branchlets, leaf blade abaxially, petioles, and inflorescences covered with gray-white tomentum; fruit ca. 1.3 cm. |
|
69 L. wilsonii |
+ |
Young branchlets, leaf blade abaxially, petioles, and inflorescences gray-brown pubescent; fruit ca. 0.8 cm. |
|
70 L. hayatae |
|
|
|
|
72 (70) |
Young branchlets densely brown tomentose; leaf blade lateral veins 10-20 pairs; umbels mostly solitary. |
|
73 L. elongata |
+ |
Young branchlets densely gray-yellow or yellow-brown villous; leaf blade lateral veins 5-10 pairs; umbels mostly clustered |
|
(73) |
|
|
|
|
73 (72) |
Leaf blade lanceolate-elliptic, narrower, less than 2.5 cm in diam., midrib slightly prominent abaxially, transverse veinlets inconspicuous abaxially. |
|
71 L. kobuskiana |
+ |
Leaf blade lanceolate, oblanceolate, or oblong-lanceolate, broader, mostly more than 2.5 cm in diam., midrib impressed adaxially, transverse veinlets conspicuously prominent. |
|
72 L. acutivena |