| 1 |
Leaves opposite or whorled (or in ser. 26, 84, 86, 87 alternate apically). |
|
(2) |
| + |
Leaves alternate or at least alternate apically (or in ser. 1, 30, 31, 33, 34, 36, 37, 45, 67, 102 rarely with pseudo-opposite or opposite leaves). |
|
(62) |
| |
|
|
|
| 2 (1) |
Leaves opposite or occasionally alternate. |
|
(3) |
| + |
Leaves usually in whorls of 3 or 4. |
|
(30) |
| |
|
|
|
| 3 (2) |
Corolla galea beakless. |
|
(4) |
| + |
Corolla galea beaked. |
|
(13) |
| |
|
|
|
| 4 (3) |
Corolla galea toothless. |
|
(5) |
| + |
Corolla galea toothed. |
|
(8) |
| |
|
|
|
| 5 (4) |
Leaves lanceolate to linear. |
|
(6) |
| + |
Leaves ± ovate. |
|
(7) |
| |
|
|
|
| 6 (5) |
Leaves entire or obscurely crenate |
|
50 Ser. Salicifoliae |
| + |
Leaves pinnatipartite to pinnatisect |
|
102 Ser. Flammeae |
| |
|
|
|
| 7 (5) |
Corolla 3--3.5 cm; roots glabrous |
|
21 Ser. Salviiflorae |
| + |
Corolla less than 2 cm; roots densely yellow-brown pubescent |
|
26 Ser. Aloenses |
| |
|
|
|
| 8 (4) |
Leaves mostly basal, stem leaves often only in 1 pair; flowers in a densely capitate spike or flowers only 2 or 3. |
|
(9) |
| + |
Leaves mostly on stem, basal leaves few; flowers not in a dense capitulum, more than 2 or 3. |
|
(10) |
| |
|
|
|
| 9 (8) |
Plants 2--4 cm tall; flowers few; galea without a protuberance at middle of margin on each side |
|
87 Ser. Collatae |
| + |
Plants usually more than 10 cm tall; galea with a protuberance at middle of margin on each side |
|
89 Ser. Cernuae |
| |
|
|
|
| 10 (8) |
Corolla galea at least as long as lower lip. |
|
(11) |
| + |
Corolla galea shorter, only ca. 1/2 as long as lower lip, apex with only 1 tooth on each side of margin. |
|
(12) |
| |
|
|
|
| 11 (10) |
Corolla apex with several teeth on each side of margin |
|
76 Ser. Lyratae |
| + |
Corolla apex obscurely 2-toothed |
|
36 Ser. Pteridifoliae |
| |
|
|
|
| 12 (10) |
Calyx relatively large, with dense reticulate veins, teeth triangular, entire |
|
38 Ser. Pentagonae |
| + |
Calyx relatively small, without dense reticulate veins, teeth leaflike, serrate |
|
84 Ser. Atrovirides |
| |
|
|
|
| 13 (3) |
Corolla tube ± elongated, more than 2 X as long as calyx. |
|
(14) |
| + |
Corolla tube not elongated, less than 2 X as long as calyx. |
|
(19) |
| |
|
|
|
| 14 (13) |
Stems usually flexuous, ascending, more than 40 cm tall. |
|
(15) |
| + |
Stems diffuse or if erect then more than 10 cm tall. |
|
(16) |
| |
|
|
|
| 15 (14) |
Corolla beak bending downward |
|
28 Ser. Flexuosae |
| + |
Corolla beak bending upward |
|
33 Ser. Omiianae |
| |
|
|
|
| 16 (14) |
Plants low; stems short or sometimes nearly absent. |
|
(17) |
| + |
Stems ± elongated, diffuse. |
|
(18) |
| |
|
|
|
| 17 (16) |
Stems short, erect, ± woody; leaf segments few; corolla beak slightly recurved |
|
75 Ser. Urceolatae |
| + |
Stems often trailing or plants ± stemless, not woody; leaves pinnatisect; corolla beak straight |
|
31 Ser. Axillares |
| |
|
|
|
| 18 (16) |
Stems herbaceous; calyx lobes shorter than calyx tube; galea not dilated apically, toothless at margin |
|
86 Ser. Sikkimenses |
| + |
Stems ± woody; calyx lobes longer than calyx tube; galea dilated apically, toothed on lower margin |
|
34 Ser. Batangenses |
| |
|
|
|
| 19 (13) |
Stems repent basally. |
|
(20) |
| + |
Stems erect, not repent if diffuse. |
|
(21) |
| |
|
|
|
| 20 (19) |
Plants annual; leaves on stem, not fernlike |
|
77 Ser. Reptantes |
| + |
Plants perennial; leaves basal, fernlike |
|
30 Ser. Vagantes |
| |
|
|
|
| 21 (19) |
Beak of galea slender, usually S-shaped; leaves narrowly lanceolate to linear-lanceolate |
|
80 Ser. Integrifoliae |
| + |
Beak of galea short, straight or curved, not S-shaped; leaves wider. |
|
(22) |
| |
|
|
|
| 22 (21) |
Corolla with 2 colors |
|
73 Ser. Binariae |
| + |
Corolla with only 1 color. |
|
(23) |
| |
|
|
|
| 23 (22) |
Leaves alternate, often pseudo-opposite throughout; flowers long pedicellate. |
|
(24) |
| + |
Leaves opposite throughout; flowers not long pedicellate. |
|
(27) |
| |
|
|
|
| 24 (23) |
Galea not crested, beak usually straight; calyx not or slightly cleft anteriorly. |
|
(25) |
| + |
Galea usually crested, sometimes inconspicuously; calyx deeply cleft anteriorly. |
|
(26) |
| |
|
|
|
| 25 (24) |
Plants 5--9 cm tall; stems 2--4; Xizang |
|
96 Ser. Mychophilae |
| + |
Plants 30--70 cm tall; stems single; not in Xizang |
|
45 Ser. Racemosae |
| |
|
|
|
| 26 (24) |
Beak curved; calyx 2- or 3-lobed |
|
97 Ser. Franchetianae |
| + |
Beak straight or falcate; calyx 3--5-lobed |
|
67 Ser. Strobilaceae |
| |
|
|
|
| 27 (23) |
Stem leaves long petiolate; inflorescences long spicate; calyx large, with dense reticulate veins, lobes entire |
|
39 Ser. Dichotomae |
| + |
Stem leaves short petiolate; inflorescences short; calyx small, without reticulate veins, lobes serrate. |
|
(28) |
| |
|
|
|
| 28 (27) |
Galea margin 1-auricled, apex of beak erose-denticulate |
|
88 Ser. Pseudoasplenifoliae |
| + |
Galea margin not auricled, apex of beak not erose-denticulate. |
|
(29) |
| |
|
|
|
| 29 (28) |
Galea marginally 2-toothed; leaves small, broadly ovate |
|
85 Ser. Remotilobae |
| + |
Galea not marginally 2-toothed; leaves larger, narrower |
|
79 Ser. Debiles |
| |
|
|
|
| 30 (2) |
Corolla galea beakless (except for a short beak in ser. 52, 22, 61, 59). |
|
(31) |
| + |
Corolla galea beaked. |
|
(45) |
| |
|
|
|
| 31 (30) |
Corolla galea toothed. |
|
(32) |
| + |
Corolla galea toothless. |
|
(36) |
| |
|
|
|
| 32 (31) |
Bases of whorled petioles of leaves or at least bracts connate, cupular |
|
40 Ser. Reges |
| + |
Bases of petioles of leaves and bracts not connate or cupular. |
|
(33) |
| |
|
|
|
| 33 (32) |
Corolla tube strongly decurved in calyx, corolla ± S-shaped |
|
55 Ser. Microchilae |
| + |
Corolla tube not strongly decurved in calyx, corolla not S-shaped. |
|
(34) |
| |
|
|
|
| 34 (33) |
Calyx lobes 2, calyx tube deeply cleft anteriorly |
|
52 Ser. Ikomanae |
| + |
Calyx lobes 5, calyx tube slightly cleft anteriorly. |
|
(35) |
| |
|
|
|
| 35 (34) |
Calyx lobes triangular or lanceolate, entire |
|
51 Ser. Rigidae |
| + |
Calyx lobes ± leaflike, serrate |
|
22 Ser. Melampyriflorae |
| |
|
|
|
| 36 (31) |
Bases of petioles of leaves and bracts connate, cupular |
|
40 Ser. Reges |
| + |
Bases of petioles of leaves and bracts not connate or cupular. |
|
(37) |
| |
|
|
|
| 37 (36) |
Corolla tube decurved basally. |
|
(38) |
| + |
Corolla tube erect or decurved apically. |
|
(39) |
| |
|
|
|
| 38 (37) |
Corolla yellow, galea plicate; anthers apiculate at apex |
|
53 Ser. Plicatae |
| + |
Corolla usually purple, red, white, or rarely yellow, galea not plicate; anthers not apiculate at apex |
|
54 Ser. Verticillatae |
| |
|
|
|
| 39 (37) |
Corolla tube ± erect. |
|
(40) |
| + |
Corolla tube decurved. |
|
(41) |
| |
|
|
|
| 40 (39) |
Plants annual; leaf segments 5--7 pairs |
|
61 Ser. Simae |
| + |
Plants perennial; leaf segments 7--17 pairs |
|
1 Ser. Gloriosae |
| |
|
|
|
| 41 (39) |
Inflorescences and calyx densely tomentose |
|
81 Ser. Pilostachyae |
| + |
Inflorescences and calyx glabrous or sparsely pubescent. |
|
(42) |
| |
|
|
|
| 42 (41) |
Corolla tube ± decurved at middle. |
|
(43) |
| + |
Corolla tube slightly curved, expanded, or bent forward apically. |
|
(44) |
| |
|
|
|
| 43 (42) |
Plants annual; stems (15--)30--80 cm tall, often branched apically; corolla 7--9 mm |
|
58 Ser. Molles |
| + |
Plants perennial; stems 5--12 cm tall, simple; corolla 22--30 mm |
|
57 Ser. Caucasicae |
| |
|
|
|
| 44 (42) |
Leaves pinnatipartite to pinnatisect; leaves whorled |
|
60 Ser. Abrotanifoliae |
| + |
Leaves pinnatilobate to pinnatifid; leaves opposite or whorled |
|
59 Ser. Denudatae |
| |
|
|
|
| 45 (30) |
Corolla tube ca. 2.5--3 X as long as calyx |
|
43 Ser. Cyathophyllae |
| + |
Corolla tube less than 3 X as long as calyx. |
|
(46) |
| |
|
|
|
| 46 (45) |
Bases of petioles of leaves and bracts connate, cupular. |
|
(47) |
| + |
Bases of petioles of leaves and bracts not connate or cupular. |
|
(48) |
| |
|
|
|
| 47 (46) |
Galea navicular; beak indistinct |
|
41 Ser. Cyathophylloides |
| + |
Galea not navicular; beak long, distinct |
|
42 Ser. Superbae |
| |
|
|
|
| 48 (46) |
Flowers scarcely 6 mm, in a dense capitate raceme |
|
25 Ser. Coniferae |
| + |
Flowers more than 10 mm. |
|
(49) |
| |
|
|
|
| 49 (48) |
Beak of galea shorter than anther-containing part. |
|
(50) |
| + |
Beak of galea longer than anther-containing part. |
|
(54) |
| |
|
|
|
| 50 (49) |
Stems with curved branches |
|
23 Ser. Longicaules |
| + |
Stems unbranched, or with erect straight branches. |
|
(51) |
| |
|
|
|
| 51 (50) |
Corolla tube decurved in calyx tube |
|
56 Ser. Cheilanthifoliae |
| + |
Corolla tube not decurved in calyx tube. |
|
(52) |
| |
|
|
|
| 52 (51) |
Corolla tube decurved in throat of calyx tube |
|
23 Ser. Longicaules |
| + |
Corolla tube not decurved or decurved outside calyx lobes. |
|
(53) |
| |
|
|
|
| 53 (52) |
Anther-containing part of galea red, goosehead-shaped; stem leaves in 1 or 2 pairs |
|
90 Ser. Tatsienenses |
| + |
Anther-containing part of galea not red, not goosehead-shaped; stem leaves numerous |
|
62 Ser. Myriophyllae |
| |
|
|
|
| 54 (49) |
Beak of galea coiled or twisted. |
|
(55) |
| + |
Beak of galea straight or slightly falcate-curved. |
|
(58) |
| |
|
|
|
| 55 (54) |
Corolla tube bent at a right angle apically |
|
83 Ser. Meteororhynchae |
| + |
Corolla tube not bent at a right angle apically. |
|
(56) |
| |
|
|
|
| 56 (55) |
Corolla yellow or purplish, with yellowish tube apically and lower lip basally |
|
74 Ser. Pectinatiformes |
| + |
Corolla purplish red or yellow, or yellow with purple galea. |
|
(57) |
| |
|
|
|
| 57 (56) |
Calyx often less than 6 mm, lobes ± serrate; bracts usually narrower and serrate or pinnatilobate |
|
63 Ser. Semitortae |
| + |
Calyx often ca. 10 mm, lobes ± entire; bracts at least ovate and entire apically |
|
64 Ser. Pectinatae |
| |
|
|
|
| 58 (54) |
Stems branched apically. |
|
(59) |
| + |
Stems unbranched apically. |
|
(60) |
| |
|
|
|
| 59 (58) |
Branches in whorls of 4--6, often woody; beak of galea slightly falcate-curved; calyx lobes ovate; capsule ca. 1/2 exceeding calyx |
|
24 Ser. Graciles |
| + |
Branches opposite or in whorls of 4, herbaceous; beak of galea bent upward; calyx lobes triangular; capsule ca. 4/5 exceeding calyx |
|
27 Ser. Moupinenses |
| |
|
|
|
| 60 (58) |
Corolla tube decurved in calyx |
|
56 Ser. Cheilanthifoliae |
| + |
Corolla tube not decurved. |
|
(61) |
| |
|
|
|
| 61 (60) |
Basal leaves large, persistent, ± lanceolate or narrowly oblong, pinnatifid, with many segments; stem leaves in whorls of 3 |
|
82 Ser. Tantalorhynchae |
| + |
Basal leaves smaller or not persistent, ± elliptic to ovate, with few segments; stem leaves sometimes opposite |
|
78 Ser. Brevifoliae |
| |
|
|
|
| 62 (1) |
Corolla slightly beaked or beakless (beak not longer than wide). |
|
(63) |
| + |
Corolla beaked. |
|
(84) |
| |
|
|
|
| 63 (62) |
Galea with marginal teeth near apex. |
|
(64) |
| + |
Galea without marginal teeth. |
|
(73) |
| |
|
|
|
| 64 (63) |
Galea with several marginal teeth on each side near apex |
|
105 Ser. Rhynchodontae |
| + |
Galea usually with a distinct marginal tooth on each side of apex. |
|
(65) |
| |
|
|
|
| 65 (64) |
Lower corolla lip usually erect or slightly spreading, corolla tube usually not decurved. |
|
(66) |
| + |
Lower corolla lip spreading, corolla tube usually decurved. |
|
(69) |
| |
|
|
|
| 66 (65) |
Plants branched |
|
44 Ser. Pedicularis |
| + |
Plants usually unbranched. |
|
(67) |
| |
|
|
|
| 67 (66) |
Basal leaves to 28 cm, with 5--9 pairs of segments |
|
36 Ser. Pteridifoliae |
| + |
Basal leaves less than 12 cm, with more segments. |
|
(68) |
| |
|
|
|
| 68 (67) |
Plants reddish or reddish brown hirtellous; basal leaves persistent |
|
20 Ser. Hirtellae |
| + |
Plants not hirtellous; basal leaves not persistent |
|
16 Ser. Striatae |
| |
|
|
|
| 69 (65) |
Leaves opposite or sometimes alternate apically; stems ascending |
|
84 Ser. Atrovirides |
| + |
Leaves alternate; stems erect. |
|
(70) |
| |
|
|
|
| 70 (69) |
Plants usually less than 15 cm tall. |
|
(71) |
| + |
Plants usually more than 15 cm tall. |
|
(72) |
| |
|
|
|
| 71 (70) |
Plants without persistent marcescent leaves at base; inflorescences more than 1/2 total height of plants, centrifugal |
|
104 Ser. Pseudo-oederianae |
| + |
Plants with persistent marcescent leaves at base; inflorescences short spicate or capitate |
|
92 Ser. Merrillianae |
| |
|
|
|
| 72 (70) |
Leaves usually 2-pinnatifid, 1-pinnatisect, 2-pinnatisect or pinnatipartite |
|
66 Ser. Comosae |
| + |
Leaves usually 1-pinnatisect, with linear-lanceolate, dentate segments |
|
65 Ser. Sudeticae |
| |
|
|
|
| 73 (63) |
Corolla tube decurved apically, lower lip spreading; capsule oblique. |
|
(74) |
| + |
Corolla tube usually erect or occasionally decurved basally, lower lip not spreading; capsule not oblique. |
|
(77) |
| |
|
|
|
| 74 (73) |
Plants usually cespitose or procumbent to ascending. |
|
(75) |
| + |
Plants erect; flowers in a spike, or few and basilar, long pedicellate. |
|
(76) |
| |
|
|
|
| 75 (74) |
Inflorescences terminal and basilar, usually distinctly centrifugal |
|
103 Ser. Corydaloides |
| + |
Flowers few, axillary |
|
87 Ser. Collatae |
| |
|
|
|
| 76 (74) |
Inflorescences short, capitate or flowers few, not distinctly centrifugal |
|
91 Ser. Roseae |
| + |
Inflorescences elongated, spicate, with dense flowers |
|
102 Ser. Flammeae |
| |
|
|
|
| 77 (73) |
Flowers axillary. |
|
(78) |
| + |
Plants with distinct erect stems; flowers usually in terminal inflorescences. |
|
(79) |
| |
|
|
|
| 78 (77) |
Plants usually stemless |
|
29 Ser. Acaules |
| + |
Plants with distinct stems |
|
26 Ser. Aloenses |
| |
|
|
|
| 79 (77) |
Apex of galea acute |
|
2 Ser. Grandiflorae |
| + |
Apex of galea usually rounded or obtuse. |
|
(80) |
| |
|
|
|
| 80 (79) |
Throat of corolla expanded, lower lip long stipitate basally |
|
3 Ser. Tsekouenses |
| + |
Throat of corolla not expanded, lower lip not stipitate. |
|
(81) |
| |
|
|
|
| 81 (80) |
Stem leaves usually few; plants not drying black |
|
1 Ser. Gloriosae |
| + |
Stem leaves usually numerous; plants drying black or not. |
|
(82) |
| |
|
|
|
| 82 (81) |
Leaves linear to linear-lanceolate, pinnatipartite; plants to 1 m tall |
|
4 Ser. Tristes |
| + |
Leaves ovate to elliptic-oblong, dentate to pinnatilobate; plants less than 40 cm tall. |
|
(83) |
| |
|
|
|
| 83 (82) |
Plants drying black; rhizomes distinct, rootstock developed, roots glabrous; leaves alternate; corolla large, galea falcate apically |
|
5 Ser. Galeatae |
| + |
Plants not drying black; without rhizomes, roots densely yellowish brown pubescent; leaves mostly opposite; corolla ca. 1.3 cm, galea ± straight |
|
26 Ser. Aloenses |
| |
|
|
|
| 84 (62) |
Corolla tube at least 2 X as long as calyx (except P. bicolor of ser. Wilsoniae). |
|
(85) |
| + |
Corolla tube less than 2 X as long as calyx. |
|
(95) |
| |
|
|
|
| 85 (84) |
Plants usually diffuse; calyx lobes usually 5. |
|
(86) |
| + |
Plants erect or ± stemless; if stems diffuse, calyx lobes 2 or 3, not 5. |
|
(88) |
| |
|
|
|
| 86 (85) |
Plants ± woody; leaves with linear segments; galea usually abruptly constricted into a straight beak |
|
34 Ser. Batangenses |
| + |
Plants herbaceous; leaves with ovate to oblong segments; galea usually gradually constricted into a beak. |
|
(87) |
| |
|
|
|
| 87 (86) |
Beak of galea bent upward |
|
33 Ser. Omiianae |
| + |
Beak of galea not bent upward, usually straight or twisted |
|
35 Ser. Muscicolae |
| |
|
|
|
| 88 (85) |
Plants more than 10 cm tall, if smaller, corolla often yellow, or corolla red-purple and less than 1.5 cm. |
|
(89) |
| + |
Plants less than 10 cm tall; corolla ± red-purple. |
|
(91) |
| |
|
|
|
| 89 (88) |
Inflorescences short, racemose, centripetal; galea 1-toothed on each side |
|
71 Ser. Amplitubae |
| + |
Inflorescences usually longer, centrifugal; galea toothless at margin. |
|
(90) |
| |
|
|
|
| 90 (89) |
Lower corolla lip spreading, not enveloping galea |
|
111 Ser. Longiflorae |
| + |
Lower corolla lip not spreading, usually enveloping galea, only slender beak projecting |
|
112 Ser. Megalanthae |
| |
|
|
|
| 91 (88) |
Pedicel to 1--4 cm; galea circinate-incurved basally |
|
101 Ser. Neolatitubae |
| + |
Pedicel shorter; galea not incurved basally. |
|
(92) |
| |
|
|
|
| 92 (91) |
Corolla tube expanded apically |
|
100 Ser. Wilsoniae |
| + |
Corolla tube not expanded apically. |
|
(93) |
| |
|
|
|
| 93 (92) |
Corolla with lower lip less than 2 cm wide; corolla tube slender, usually less than 0.75 mm wide |
|
99 Ser. Albiflorae |
| + |
Corolla with lower lip more than 2 cm wide; corolla tube usually more than 1 mm wide. |
|
(94) |
| |
|
|
|
| 94 (93) |
Leaves pinnatifid; beak of galea usually downward uncinate |
|
108 Ser. Robustae |
| + |
Leaves ± entire or only pinnatifid; beak of galea not downward uncinate |
|
110 Ser. Pumiliones |
| |
|
|
|
| 95 (84) |
Galea usually marginally long ciliate. |
|
(96) |
| + |
Galea not marginally long ciliate. |
|
(108) |
| |
|
|
|
| 96 (95) |
Galea navicular. |
|
(97) |
| + |
Galea not navicular. |
|
(100) |
| |
|
|
|
| 97 (96) |
Leaves linear to linear-lanceolate or oblong-lanceolate, pinnatilobate or dentate |
|
8 Ser. Craspedotrichae |
| + |
Leaves not linear, shorter and wider, segmented. |
|
(98) |
| |
|
|
|
| 98 (97) |
Leaves pinnatisect, with linear-lanceolate segments |
|
17 Ser. Proboscideae |
| + |
Leaves double dentate to pinnatipartite, with shorter wider segments. |
|
(99) |
| |
|
|
|
| 99 (98) |
Leaves ± lanceolate-oblong, less than 3.5 X as long as wide |
|
7 Ser. Dolichocymbae |
| + |
Leaves not linear to linear-lanceolate, more than 4 X as long as wide |
|
6 Ser. Rudes |
| |
|
|
|
| 100 (96) |
Beak of galea nearly as long as lower corolla lip. |
|
(101) |
| + |
Beak of galea longer than lower corolla lip. |
|
(104) |
| |
|
|
|
| 101 (100) |
Leaves mostly basal, pinnatisect with lanceolate segments; beak of galea with apical tuft of hairs |
|
18 Ser. Lachnoglossae |
| + |
Leaves mostly on stem, not pinnatisect; beak of galea without apical tuft of hairs. |
|
(102) |
| |
|
|
|
| 102 (101) |
Lobes of lower corolla lip long stipitate basally, galea ± abruptly constricted into a slender beak |
|
11 Ser. Lasiophrydes |
| + |
Lobes of lower corolla lip not stipitate, galea gradually constricted into a beak. |
|
(103) |
| |
|
|
|
| 103 (102) |
Plants usually more than 30 cm tall, to 80 cm; corolla galea less than 3 mm wide, sparsely pubescent |
|
9 Ser. Imbricatae |
| + |
Plants usually less than 30 cm tall; corolla galea 4--5 mm wide, densely pubescent |
|
10 Ser. Trichoglossae |
| |
|
|
|
| 104 (100) |
Leaves linear or linear-lanceolate. |
|
(105) |
| + |
Leaves not narrowly linear. |
|
(106) |
| |
|
|
|
| 105 (104) |
Plants usually more than 30 cm tall, to 1 m; beak of galea not twisted; lobes of lower corolla lip narrowly lanceolate |
|
12 Ser. Kongboenses |
| + |
Plants less than 30 cm tall; beak of galea S-shaped; lobes of lower corolla lip short, wide |
|
15 Ser. Kialenses |
| |
|
|
|
| 106 (104) |
Leaves ± ovate, usually more than 3.5 cm wide |
|
14 Ser. Excelsae |
| + |
Leaves linear-lanceolate to lanceolate-oblong, usually less than 3.5 cm wide (except P. angustilabris to 5.5 cm wide). |
|
(107) |
| |
|
|
|
| 107 (106) |
Galea not twisted, densely villous at base and at margin; lobes of lower corolla lip not square-shaped |
|
19 Ser. Recurvae |
| + |
Galea strongly twisted, glabrous or villous on lower margin and beak; lobes of lower corolla lip nearly square-shaped |
|
15 Ser. Kialenses |
| |
|
|
|
| 108 (95) |
Inflorescences distinctly centrifugal. |
|
(109) |
| + |
Inflorescences distinctly centripetal or indistinctly centrifugal. |
|
(111) |
| |
|
|
|
| 109 (108) |
Beak of galea more than 3 X as long as lower corolla lip |
|
107 Ser. Macrorhynchae |
| + |
Beak of galea shorter or slightly longer than lower corolla lip. |
|
(110) |
| |
|
|
|
| 110 (109) |
Lower corolla lip less than 1.2 cm wide |
|
106 Ser. Filiculae |
| + |
Lower corolla lip usually more than 1.5 cm wide |
|
109 Ser. Macranthae |
| |
|
|
|
| 111 (108) |
Stems weak, slender, usually diffuse; flowers usually axillary. |
|
(112) |
| + |
Stems erect or ± ascending, or ± stemless; flowers arising from a stock. |
|
(115) |
| |
|
|
|
| 112 (111) |
Flowers usually in clusters at base of stems |
|
30 Ser. Vagantes |
| + |
Flowers not in clusters at base of stems, solitary. |
|
(113) |
| |
|
|
|
| 113 (112) |
Galea with pairs of marginal teeth |
|
94 Ser. Odontophorae |
| + |
Galea without marginal teeth. |
|
(114) |
| |
|
|
|
| 114 (113) |
Leaves without distinct reticulate veins abaxially; calyx tube without reticulate veins |
|
31 Ser. Axillares |
| + |
Leaves with distinct reticulate veins abaxially; calyx tube with reticulate veins apically |
|
32 Ser. Infirmae |
| |
|
|
|
| 115 (111) |
Inflorescences usually elongated, not interrupted. |
|
(116) |
| + |
Inflorescences usually subcapitate, few flowered, or ± elongated and interrupted basally. |
|
(117) |
| |
|
|
|
| 116 (115) |
Stems to 80 cm tall; leaves relatively large; corolla relatively small, beak of galea bent upward, lower lip 1/4 to 1/3 as long as galea |
|
13 Ser. Subsurrectae |
| + |
Stems shorter; leaves relatively small; corolla relatively large, beak of galea bent downward only, lower lip longer |
|
69 Ser. Oxycarpae |
| |
|
|
|
| 117 (115) |
Plants less than 10 cm tall, without a distinct main stem. |
|
(118) |
| + |
Plants more than 10 cm tall. |
|
(121) |
| |
|
|
|
| 118 (117) |
Leaves numerous, persistent, with several narrow segments |
|
93 Ser. Asplenifoliae |
| + |
Leaves few, not persistent, with fewer wider segments. |
|
(119) |
| |
|
|
|
| 119 (118) |
Lower corolla lip more than 2 cm wide, galea usually robust, apex of beak 2-cleft |
|
108 Ser. Robustae |
| + |
Lower corolla lip narrower, galea narrower, apex of beak not 2-cleft. |
|
(120) |
| |
|
|
|
| 120 (119) |
Stems rigid; leaves alternate; roots numerous |
|
95 Ser. Paucifoliatae |
| + |
Stems usually weak, flexuous; leaves usually pseudo-opposite; roots few |
|
96 Ser. Mychophilae |
| |
|
|
|
| 121 (117) |
Pedicel to 3--11.5 cm; galea not twisted |
|
97 Ser. Franchetianae |
| + |
Pedicel less than 3 cm; galea strongly twisted laterally. |
|
(122) |
| |
|
|
|
| 122 (121) |
Flowers ± in a capitulum. |
|
(123) |
| + |
Flowers ± in a raceme. |
|
(128) |
| |
|
|
|
| 123 (122) |
Corolla more than 3 cm; lower lip relatively wide, often completely enveloping galea |
|
98 Ser. Pseudomacranthae |
| + |
Corolla usually less than 2.5 cm; lower lip relatively narrow, often spreading, not enveloping galea. |
|
(124) |
| |
|
|
|
| 124 (123) |
Leaves to 2.5--13 X 14 cm, stem leaves often ovate-elliptic |
|
37 Ser. Phaceliifoliae |
| + |
Leaves smaller, ± oblong or lanceolate. |
|
(125) |
| |
|
|
|
| 125 (124) |
Plants usually branched basally and at middle |
|
67 Ser. Strobilaceae |
| + |
Plants unbranched or only branched basally. |
|
(126) |
| |
|
|
|
| 126 (125) |
Corolla to 2.8 cm; beak of galea very long, S-shaped or semicircular |
|
72 Ser. Rhinanthoides |
| + |
Corolla smaller; beak of galea shorter, not S-shaped or semicircular. |
|
(127) |
| |
|
|
|
| 127 (126) |
Leaves pinnatifid or pinnatipartite |
|
95 Ser. Paucifoliatae |
| + |
Leaves pinnatisect or pinnatipartite |
|
68 Ser. Oliganthae |
| |
|
|
|
| 128 (122) |
Galea strongly twisted laterally at base |
|
70 Ser. Dissectae |
| + |
Galea not strongly twisted laterally at base. |
|
(129) |
| |
|
|
|
| 129 (128) |
Leaves often ovate or elliptic, with few segments |
|
49 Ser. Furfurae |
| + |
Leaves oblong to linear, with numerous segments, some merely crenate. |
|
(130) |
| |
|
|
|
| 130 (129) |
Basal leaves cespitose; plants herbaceous; bracts petiolate |
|
95 Ser. Paucifoliatae |
| + |
Basal leaves not cespitose; plants ± woody; bracts sessile. |
|
(131) |
| |
|
|
|
| 131 (130) |
Leaves only double dentate or entire. |
|
(132) |
| + |
Leaves pinnatipartite to pinnatisect. |
|
(133) |
| |
|
|
|
| 132 (131) |
Beak of galea distinct |
|
45 Ser. Racemosae |
| + |
Beak of galea indistinct |
|
46 Ser. Carnosae |
| |
|
|
|
| 133 (131) |
Pedicel relatively short; beak of galea short or obscure |
|
47 Ser. Microphyllae |
| + |
Pedicel relatively long, distinct; beak of galea slender |
|
48 Ser. Polyphyllatae |