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11. Scutellaria Linnaeus, Sp. Pl. 2: 598. 1753.

黄芩属 huang qin shu

Scutellaria hypericifolia

Credit: Harvard University Herbaria

Herbs or subshrubs, rarely shrubs, not aromatic. Leaves entire to pinnatifid. Inflorescences terminal or axillary racemes or spikes; floral leaves usually bractlike apically. Flowers axillary, opposite or sometimes alternate apically. Calyx short tubular, dorsiventrally flattened, 2-lipped; lips entire, closed and ultimately divided to base along sutures in fruit; upper lip deciduous, with a transverse, rounded, concave, scalelike scutellum (shield) or without and abaxially conspicuously saccate; lower lip persistent. Corolla 2-lipped; tube exserted, arcuate or suberect, gradually widening to throat, base bent and saccate or spurred, usually puberulent annulate outside; upper lip erect, galeate; lower lip 3-lobed, middle lobe broad, flattened, entire, lateral lobes ± joined to upper lip and sometimes spreading. Stamens 4, didynamous, anterior 2 longest, underlying upper lip; anthers close together in pairs, bearded on cell aperture; posterior pair conspicuously 2-celled, ± apically acute, anterior pair 1-celled by abortion. Style subulate, apically unequally 2-cleft. Nutlets oblate, globose, to ovoid.

About 350 species: worldwide, but only a few in tropical Africa; 98 species in China.

A very isolated genus with unsatisfactory traditional divisions. Paton (Kew Bull. 45: 399-450. 1990) has proposed a revised classification of Scutellaria but did not deal with a large number of the Chinese taxa, and it is not possible to utilize his divisions in detail.


Key 7

1 Leaf blade abaxially white or gray tomentose.   (2)
+ Leaf blade abaxially green or gray-green, sparsely or densely erect or floccose-pubescent, sometimes with intermixed glandular hairs.   (3)
       
2 (1) Leaves pinnatipartite   90 Scutellaria przewalskii
+ Leaves pinnately lobed, margin incised-serrate   89 Scutellaria sieversii
       
3 (1) Leaf serrations conspicuous, spreading.   (4)
+ Leaf serrations inconspicuous, always coalescent.   (5)
       
4 (3) Corolla yellow, without purple spots   91 Scutellaria supina
+ Corolla yellowish, with conspicuous purple spots   92 Scutellaria prostrata
       
5 (3) Bracts pubescent and glandular along veins and margin, otherwise subglabrous   94 Scutellaria altaica
+ Bracts spreading pilose and glandular along veins and margin.   (6)
       
6 (5) Stems densely spreading pilose and pubescent throughout; petiole 0.5-1.7 cm; lower floral leaves serrate   95 Scutellaria orthotricha
+ Stems ± spreading pilose and glandular apically; petiole 0.2-0.5 cm; lower floral leaves entire   93 Scutellaria oligodonta

List of Keys

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