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Viola L., Sp. Pl., 933. 1753. Gen. Pl. ed. 5.,402, 1754; W. Becker in Beih. Bot. Centralbl. Abt. 2, 34:208, 1916; ibid. 373, 1917; in Bot. Jahrb. 54 Beibl. 120:156, 1917; in Beih. Bot. Centralbl. Abt. 2, 36:15, 1918; ibid. 40:20, 69, 119, 1923; in Engl. & Prant. Pfl. Fam. ed. 2, 21:363, 1925; Nakai in Bot. Mag. Tokyo 30:276, 1916; ibid. 36:52, 84, 118, 1922; in Bull. Soc. Bot. France 72:180, 1925; Lin in Taiwania 1:269, 1950; Maekawa in Hara Enum. Pl. Jap. 194 1954; Hashimoto in Bull. Jap. Assoc. Bot. Gard. 1959:29; M. Jacob & D. M. Moore in van Steenis, Fl. Malesiana 7:198, 1971; Hsieh in Li et al., Fl. Taiwan 3:769, 1977.
菫菜屬
Perennial herb. Leaves alternate, petiolate; blade suborbicular to linear-lanceolate, the margin serrate to crenate. Stipules free or adnate to petiole, persistent, usually fimbriate or serrate, sometimes nearly entire. Flowers axillary, bisexual, zygomorphic, solitary, with a pair of bracteoles in the half of the peduncles; sepals 5, nearly equal, entire to denticulate, with appendages at base, persistent; petals 5, unequal, the lower one spurred at base, the lateral pair often bearded inside. Stamens 5, connivent around the gynoecium; anther 2-celled, subsessile, connective produced into an apical appendage, the lower two bearing appendages projecting into the spur. Ovary 1-celled, glabrous or pubescent, ovules numerous on 3-parietal placentas; style straight or geniculate at base, filiform to conspicuously clavate, often lobed or more or less dilated at apex, with a openning of stigmatic cavity terminal or anterior and subterminal. Fruit a 3-valved capsule globose to ellipsoidal; valves boat-shaped usually with thick rigid keels. Seeds numerous, ovoid or ellipsoidal with crustaceous testa.
More than 400 species chiefly in temperate regions throughout the world. 18 taxa and one doubtful species in Taiwan.
Hashimoto, T. 1987. Trigonocarpae violets of Taiwan. Ann. Tsukuba Bot. Gard. 6: 1-7; Wang, J. C. & T. C. Huang 1990. Note on the flora of Taiwan (9). The genus Viola L. (Violaceae). Taiwania 35: 12-56.
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1 |
Flower yellow; leaves reniform to broadly ovate, apex rounded or obtuse |
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Viola biflora |
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Flower purple or white |
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(2) |
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2 (1) |
Plants without stems or stolons; stipule adnate to petiole more than 1/2 of length |
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(3) |
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Plants with stems or stolons |
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(7) |
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3 (2) |
Stipule adnate to petiole, less than 2/3 of length; basal petal rounded at apex; rhizome long creeping, nodose; leaves triangular-ovate |
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Viola senzanensis |
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Stipule adnate to petiole, up to 3/4 of length; basal petal truncate or emarginate at apex; rhizome short, stout |
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(4) |
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4 (3) |
Roots dark brown |
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Viola mandshurica |
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Roots white or yellowish brown |
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(5) |
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5 (4) |
Spur slender, 2.5-6 mm long, lateral petals usually beardless |
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Viola confusa |
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Spur short, up to 3.5 mm long, lateral petals bearded |
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(6) |
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6 (5) |
Upper petals usually bearded |
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Viola betonicifolia |
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Upper petals beardless |
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Viola inconspicua ssp. nagasakiensis |
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7 (2) |
Petioles winged; basal petal shortest, acute or acicular at apex |
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(8) |
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Petiole wingless |
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(11) |
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8 (7) |
Flower less than 1.5 cm across; plants with stems and stolons |
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(9) |
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Flower more than 1.5 cm in diameter; plants without stems |
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(10) |
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9 (8) |
leaves cuneate or shallowly cordate at base; lateral petals usually bearded |
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Viola diffusa |
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Leaves cordate at base; lateral petals usually beard-less |
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Viola tenuis |
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10 (8) |
Leaves ovate, lateral petals beardless |
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Viola nagasawai var. nagasawai |
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Leaves triangular-ovate, lateral petals bearded |
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Viola nagasawai var. pricei |
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11 (7) |
Basal petal truncate or emarginate at apex |
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(12) |
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Basal petal rounded or obtuse at apex |
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(16) |
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12 (11) |
Flowers small less than 1.5 cm in diameter |
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Viola arcuata |
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Flowers large, more than 2 cm in diameter |
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(13) |
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13 (12) |
Spur slender, more than 4 mm long; basal petal longest, emarginate at apex |
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(14) |
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Spur rounded, less than 2 mm long, basal petal shortest, truncate or emarginate at apex |
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(15) |
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14 (13) |
Leaves orbicular or orbicular-cordate, rounded or obtuse at apex |
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Viola formosana var. formosana |
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Leaves cordate, acute at apex |
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Viola formosana var. stenopetala |
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15 (13) |
Leaves membraneous, usually hirsute; lateral petals usually bearded |
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Viola adenothrix var. adenothrix |
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Leaves chartaceous to subcoriaceous, usually glabrous; lateral petals usually beardless |
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Viola adenothrix var. tsugitakaensis |
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16 (11) |
Lateral petals bearded; style papillose at apex; capsule globose |
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Viola shinchikuensis |
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Lateral petals beardless; style smooth at apex; capsule ellipsoid |
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(17) |
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17 (16) |
Leaf-blades cordate, usually as long as or shorter than wide |
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Viola grypoceras |
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Leaf-blades triangular ovate, usually longer than wide |
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Viola obtusa var. tsuifengensis |
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List of lower taxa
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